Новые докодонты из средней юры Сибири и новый анализ филогенетических связей докодонтов

А.О. Аверьянов, А.В. Лопатин, С.А. Краснолуцкий и С.В. Иванцов

Труды Зоологического института РАН, 2010, 314(2): 121–148 · https://doi.org/10.31610/trudyzin/2010.314.2.121

Резюме

Два новых докодонта, Simpsonodon sibiricus sp. nov. и Hutegotherium yaomingi gen. et sp. nov., и новый экземпляр Itatodon tatarinovi Lopatin et Averianov, 2005 описаны из среднеюрской (бат) итатской свиты в Красноярском крае, Западная Сибирь, Россия. Согласно новому филогенетическому анализу, включающему 37 признаков зубов и нижней челюсти и 18 таксонов докодонтов и внешней группы, Itatodon является наиболее базальным из известных докодонтов. Другие докодонты классифицированы в Docodontidae (Docodon и Haldanodon; Borealestes является стволовым докодонтидом), Simpsonodontidae fam. nov. (Simpsonodon и Dsungarodon), Tegotheriidae (Krusatodon, Hutegotherium gen. nov., Tegotherium и Sibirotherium), и Docodonta incertae sedis (Castorocauda, Tashkumyrodon и Acuoduolodon). Simpsonodontidae fam. nov. характеризуются, в частности, сильно развитой дополнительной бороздой дорсальнее меккелевой борозды. Среднеюрские млекопитающие Евразии теперь известны по трем основным фаунам: Форест-Марбл в Англии, балабансайской в Киргизии и итатской в Западной Сибири. Все эти фауны очень похожи и включают докодонтов, аллотериев, эутриконодонтов, «симметродонтов», дриолестид и стволовых затериев. Среднеюрская радиация млекопитающих практически не имеет корней в известной фауне млекопитающих ранней юры. Эволюционные события, приведшие к формированию основных групп мезозойских млекопитающих (Docodonta, Multituberculata, Eutriconodonta, Dryolestida и стволовые Zatheria), должны были произойти в интервале между синемюром и байосом в Лавразии, вероятнее всего в Азии (Сибири?).

Ключевые слова

Docodonta, Mammalia, средняя юра, Россия, Западная Сибирь

Поступила в редакцию 12 января 2010 г. · Принята в печать 11 февраля 2010 г. · Опубликована 25 июня 2010 г.

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