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Genus Hadeocoleus Ponomarenko, 1969 (Myxophaga: Schizophoridae): atlas of extinct beetle taxa

A.G.Kirejtshuk & А.G. Pоnоmаrеnkо


 

Genus Hadeocoleus Ponomarenko, 1969 (type species: Hаdeocoleus gigas Ponomarenko, 1969, monotype)

The genus includes three species of large body size (12-40 mm) with widely explanate sides of pronotum and long elytra, moderately long and thin hind legs, especially tibiae and tarsi, as well as moderately large metacoxa. Outwardly, and especially the structure of the prothorax this genus is similar to the genus Hadeocoleodes, but differs from the latter in the larger size of body, long prosternal process and metaventrite less narrowed anteriorly.

 

Composition:

 

+ Hаdeocoleus catachtonius Ponomarenko, 1969 (Lower Triassic)

+ Hаdeocoleus gigas Ponomarenko, 1969 (Lower Triassic)

+ Hаdeocoleus pelopius Ponomarenko, 1969 (Lower Triassic)

 

 


Hadeocoleus gigas Ponomarenko, 1969, holotypus («2240/322», collection PIN);

Body, from above; Lower Triassic, Middle Asia, Djailaucho).
Photo by authors, made with the assistance of A. Mazin (PIN).

Hadeocoleus gigas Ponomarenko, 1969, holotypus («2240/322», collection PIN);

Body, from below; Lower Triassic, Middle Asia, Djailaucho).
Photo by authors, made with the assistance of A. Mazin (PIN).



 

 

Literature:

 

(See also the page Systematic list of fossil beetles of Cupedina, Carabina and Scarabaeina (1 part) )

 

Пономаренко А.Г. 1969. Историческое развитие жесткокрылых-архостемат. (Труды Палеонтологического института АH СССР. 125) Москва. Наука. 1-239.

 

Пономаренко А.Г., Жерихин В.В. 1980. Надотряд Scarabaeidea. Жесткокрылообразные. (Труды Палеонтологического института АH СССР. 175) Москва, Наука. 75-84.

 

Crowson R.A. 1975. The evolutionary history of Coleoptera as documented by fossil and comparative evidence. Atti Congr. naz. Ital. Entomol. 10th. Sassari 1974. 47-90.

 

Crowson R.A. 1981. The Biology of the Coleoptera. London-New York-Toronto-Sydney-San Francisco, Academic press. 802 p.

 

Kirejtshuk A.G., 1991/1992. Evolution of mode of life as the basis for division of the beetles into groups of high taxonomic rank. M. Zunino, X. Belles, M. Blas, eds. Advances in Coleopterology. Baecelona, Ass. Europ. Coleopt. P. 249-261.

 

Ponomarenko A.G. 2002. 2.2.1.3.2 Superorder Scarabaeidea Laicharting, 1781, Order Coleoptera Linne, 1758. The beetles. In: Rasnitsyn, A.P. and Quicke, D.L.J. (eds). 2002. History of insects. Kluwer Academic Publishers, Dordrecht, Boston, London: 164-180.