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Иллюстрация из: Clarke H. Scholtz, Vasily V. Grebennikov. 2005. Scarabaeoidea Latreille, 1802. Глава из книги: Handbook of Zoology. Arthropoda. Insecta. Coleoptera. Volume 1. Coleoptera, Beetles. Morphology and Systematics. Ed. by Rolf G. Beutel, Richard A.B. Leschen. Published by Walter de Gruyter GmbH & Co KG. Copyright. Page 368.
