S.V. Kazantsev: "Morphology of the family Lycidae and some ideas on eolution of Coleoptera" (2003)

Sergey V. Kazantsev

MORPHOLOGY OF LYCIDAE WITH SOME CONSIDERATIONS ON EVOLUTION OF THE COLEOPTERA
ELYTRON, 2003, VOL. 17: 49-226
(ISSN: 0214-1353)

ABSTRACT
Morphology of Lycidae with some considerations on evolution of the Coleoptera. External and internal morphology of the family Lycidae and allied taxa is studied and analyzed. A cladistic analysis of the group is carried out. An evolution scenario of the lineage is discussed and a phylogenetic tree and a key to subfamilies are proposed. Dexoridae Kleine stat. n. is raised to the family level. Lyropaeinae Bocсk & Bocсkovс stat. n. and Miniduliticolinae Kazantsev stat. n. are raised to the subfamily level within the Lycidae. Thilmanus Baudi is transferred from Drilidae to Thilmaninae subfam. n. within the Lycidae. Platerodrilini tr. n. is described. The Lycidae are hypothesized to consist of seven subfamilies, Lyropaeinae stat. n., Leptolycinae, Ateliinae, Thilmaninae, subfam. n., Miniduliticolinae, stat. n., Lycinae and Calochrominae. Lyponiina Bocak & Bocakova is transferred from Platerodini to Calochrominae incertae sedis. Phylogeny of the Coleoptera is discussed. The Coleoptera is hypothesized to be a polyphyletic taxon, not including the common ancestor of its two major lineages, the Polyphaga and the Archostemata-Adephaga-Myxophaga.
Key words: Coleoptera, Lycidae, morphology, phylogeny, taxonomy.
 

• INTRODUCTION
• MATERIAL AND METHODS
• MORPHOLOGY
• IMMATURES STAGES
• EGG/EMBRYO
• LARVAE 1

• LARVAE 2
• PUPA
• ADULTS 1
  • Cuticle
  • Setation
  • Head 1

• ADULTS 2
  • Head 2
  • Cervix
  • Prothorax
  • Pronotum
  • Prosternum
  • Mesothorax
  • Mesopleuron
  • Mesoventrite
  • Elytron

• ADULTS 3
  • Metathorax
  • Metanotum
  • Metapleuron
  • Metaventrite
  • Metendosternite
  • Thoracic spiracles
  • Metathoracic wings 1

• ADULTS 4
  • Metathoracic wings 2
  • Articulation of the wing
  • Leg
  • Pro- and mesotrochantin
  • Metatrochantin
  • Pro- and Mesocoxae
  • Metacoxae
  • Trochanters
  • Femora and Tibiae
  • Tarsi
  • Claws
  • Abdomen
  • Tergites
  • Sternites
  • Spiculum gastrale
  • Spiculum ventrale
  • Abdominal spiracles
  • Paraproct
  • Proctiger
  • External male genitalia
  • External female genitalia
  • Internal female genitalia
• BIOLOGY
• PHYLOGENY 1

• PHYLOGENY 2
• The analysis of the data matrix of 68 characters
• NEW TAXA
  • Dexoridae Kleine, 1933, stat. n. (type genus: Dexoris Waterhouse, 1878)
  • Lyropaeinae Bocсk & Bocсkovс, 1989, stat. n. (type genus: Lyropaeus Waterhouse, 1878)
  • Thilmaninae subfam. n. (type genus: Thilmanus Baudi, 1871)
  • Miniduliticolinae stat. n. (type genus: Miniduliticola Kazantsev, 2002)
  • Platerodrilini tr. n. (type genus: Platerodrilus Pic, 1921)
• KEY TO SUBFAMILIES OF LYCIDAE
• SOME CONSIDERATIONS ON EVOLUTION OF THE COLEOPTERA
  • The Strepsiptera, not so long ago considered a family within the Coleoptera (Stylopidae, e.g. in Crowson, 1981) has proved to be the sister group of Diptera (Wheeler et al., 2001).
  • The Coleoptera sensu lato has thus already been demonstrated to be a polyphyletic taxon.
• ACKNOWLEDGEMENTS
• REFERENCES

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