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Ainsley Seago, Jose Adriano Giorgi, Jiahui Li, Adam Slipinski. 2011.
Phylogeny, classification and evolution of ladybird beetles (Coleoptera: Coccinellidae) based on simultaneous analysis of molecular and morphological data.
Molecular Phylogenetics and Evolution, 2011, 60: 137-151.
doi:10.1016/j.ympev.2011.03.015