Classification of extinct and recent Coleoptera

(besides the author and year for most taxa
the codes of geochronologic divisions and marks
on distribution in the Palaearctic region and Russia are included)

 Ordo: COLEOPTERA Linnaeus [ ... ] Pal/Rus
 =  Scarabaeida Laicharting, 1781
+Subordo: Protocoleoptera [ ... ]
 =  Archecoleoptera
+Superfamilia: Tshekardocoleoidea Rohdendorf, 1944 [ ... ] Pal/Rus
+Familia: Tshekardocoleidae Rohdendorf, 1944 [P1ar ... P2kz] Pal/Rus
+Familia: Labradorocoleidae Ponomarenko, 1969 [Kal3-cm1 ... Kal3-cm1]
+Superfamilia: Permocupedoidea Martynov, 1933 [P2uf ... T3k] Pal/Rus
+Familia: Permocupedidae Martynov, 1933 [P2uf ... P2t] Pal/Rus
 =  Kaltanocoleidae Rohdendorf, 1961
+Familia: Taldycupedidae Rohdendorf, 1961 [P2kz ... T3k] Pal/Rus
 Subordo: Archostemata Kolbe, 1908 [ ... ] Pal/Rus
 =  Cupedina
 Infraordo: Cupediformia {Kirejtshuk, 1991} [ ... ] Pal/Rus
 =  Cupedomorpha ?
 Superfamilia: Cupedoidea Laporte, 1836 [ ... ] Pal/Rus
 Familia: Cupedidae Laporte, 1836 [T21 ... R] Pal/Rus
 =  Brochocoleidae Hong, 1982
 =  Cupesidae Lacordaire, 1858
 Familia: Crowsoniellidae Iablokoff-Khnzorian, 1983 [ ... R] Pal/-
 Familia: Ommatidae Sharp et Muir, 1912 [J1he ... R] Pal/Rus
 =  Ommadidae
 =  Tetraphaleridae
 Subfamilia: Ommatinae Sharp et Muir, 1912 [??? ... R] Pal/Rus
 Subfamilia: Tetraphalerinae Crowson, 1962 [??? ... R] Pal/Rus
+Familia: Triadocupedidae Ponomarenko, 1966 [T2-3l-c ... ???] Pal/?
 Familia: Jurodidae Ponomarenko, 1985 [J12 ... R] Pal/Rus
 =  Sikhotealiniidae Lafer, 1996
 Infraordo: Micromaltiformia Kirejtshuk, 1991 [ ... R] Pal/-
 Superfamilia: Micromalthoidea ?????? [K1ap, Leba ... R] Pal/-
 Familia: Micromalthidae Barber, 1913 [K1ap, Leba ... R] Pal/-
 Subordo: INCERTAE SEDIS: [ ... ]
+Superfamilia: Ademosynoidea Ponomarenko, 1969 [ ... ]
+Familia: Ademosynidae Ponomarenko, 1969 [P2t ... K12] Pal/?
 Subordo: Polyphaga Emery, 1886 [ ... ] Pal/Rus
 =  Scarabaeida
 Infraordo: Staphyliniformia Lameere, 1900 [ ... ] Pal/Rus
 =  Scarabaeiformia ???
 =  Scarabaeomorpha
 =  Staphylinomorpha
 Superfamilia: Hydrophyloidea Latreille, 1802 [ ... ] Pal/Rus
 Familia: Hydraenidae Mulsant, 1844 [J11 ... R] Pal/Rus
 =  Limnebiidae Mulsant, 1844
 =  Ochtebiidae C.G. Thomson, 1859
 Subfamilia: Prostethopinae Perkins in Perkins et Balfour-Browne, 1994 [ ... R] ?/?
 Subfamilia: Hydraeninae Mulsant, 1844 [ ... R] Pal/Rus
 Subfamilia: Ochthebiinae C.G. Thomson, 1859 [Pg1d ... R] Pal/Rus
 Familia: Hydrochidae C.G. Thomson, 1859 [Pg22 ... R] Pal/Rus
 Familia: Helophoridae Leach, 1815 [N13 ... R] Pal/Rus
 =  Elophorinae
 Familia: Spercheidae Erichson, 1837 [ ... R] Pal/Rus
 Familia: Hydrophilidae Latreille, 1802 [J11 ... R] Pal/Rus
 =  Sphaeridiidae Latreille, 1802
 Subfamilia: Epimetopinae Zaitzev, 1908 [ ... R] ?/?
 Subfamilia: Horelophinae Hansen, 1991 [ ... R] -/-
 Subfamilia: Sphaeridiinae Latreille, 1802 [ ... R] Pal/Rus
 Subfamilia: Hydrobiinae Mulsant, 1844 [ ... R] Pal/Rus
 Subfamilia: Chetarthriinae [ ... R] ?/?
 Subfamilia: Horelophinae Hansen, 1991 [ ... R] Pal/Rus
 Subfamilia: Hydrophilinae Latreille, 1802 [ ... R] Pal/Rus
 =  Amphiopinae Orchym., 1919
 Subfamilia: Berosinae Mulsant, 1844 [Pg22, GreR ... R] Pal/Rus
 Familia: Georissidae Laporte, 1819 [Pg22 ... R] Pal/Rus
 =  Georyssidae Laporte, 1819
 Superfamilia: Histeroidea Gyllenhal, 1808 [ ... ] Pal/Rus
 Familia: Sphaeritidae Shuckard, 1839 [ ... R] Pal/Rus
 Familia: Synteliidae Lewis, 1882 [ ... R] Pal/Rus
 Familia: Histeridae Gyllenhal, 1808 [Pg22 ... R] Pal/Rus
 =  Niponiidae Handlirsch, 1925
 Subfamilia: Dendrophilinae Reitter, 1909 [Pg22, Mess ... R] Pal/Rus
 Subfamilia: Hetaeriinae Marseul, 1857 [ ... R] Pal/Rus
 Subfamilia: Histerinae Gyllenhal, 1808 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Onthophilinae MacLeay, 1819 [ ... R] Pal/Rus
 Subfamilia: Tribalinae Bickhardt, 1914 [ ... R] Pal/Rus
 Subfamilia: Abraeinae MacLeay, 1819 [ ... R] Pal/Rus
 Subfamilia: Chlamydopsinae Bickhardt, 1914 [ ... R] ?/?
 Subfamilia: Niponiinae Fowler, 1912 [ ... R] ?/?
 Subfamilia: Saprininae Blanchard, 1845 [ ... R] Pal/Rus
 Subfamilia: Trypanaeinae Marseul, 1857 [ ... R] ?/?
 =  Triponaeinae
 Subfamilia: Trypeticinae Bickhardt, 1914 [ ... R] ?/?
 Superfamilia: Staphylinoidea Latreille, 1802 [ ... ] Pal/Rus
 Familia: Agyrtidae C.G. Thomson, 1859 [J12 ... R] Pal/Rus
 =  Silphidae, part.
 Subfamilia: Lyrosominae Horn, 1880 [ ... R] ?/?
 Subfamilia: Pterolominae Port., 1914 [ ... R] Pal/Rus
 Subfamilia: Catopocerinae Hatch, 1927 [ ... R] ?/?
 =  Glacicavicolinae
 =  Pinodytinae Horn, 1880
 Familia: Colonidae Horn, 1880 [ ... R] Pal/Rus
 =  Leptinidae LeConte, 1866
 =  Platypsilidae Ritsema, 1869
 Subfamilia: Coloninae Horn, 1880 [ ... ] Pal/Rus
 Subfamilia: Platypsilinae Ritsema, 1869 [ ... R] Pal/Rus
 Subfamilia: Leptininae LeConte, 1866 [ ... R] Pal/Rus
 Familia: Leiodidae Fleming, 1821 [K12 ... R] Pal/Rus
 =  Agathidiidae Westwood, 1839
 =  Anisotomidae Erichson, 1845
 =  Camiaridae Jeannel, 1911
 =  Catopidae Jeannel, 1922
 =  Cholevidae Jeannel, 1922
 =  Leptodiridae Abeille de Perrin, 1878
 =  Liodidae Jacobson, 1910
 =  Sogdiidae
 Subfamilia: Camiarinae Jeannel, 1911 [ ... R] ?/?
 =  Agyrtodinae
 Subfamilia: Liodinae Fleming, 1951 [ ... R] Pal/Rus
 =  Anisotominae
 Subfamilia: Cholevinae Kirby, 1837 [K12 ... R] Pal/Rus
 =  Anemadinae Jeannel, 1922
 =  Bathysciinae Horn, 1880
 =  Catopinae Jeannel, 1922
 =  Eucatopinae Jeannel, 1921
 =  Leptodirinae Abeille de Perrin, 1878
 =  Nemadinae
 =  Ptomaphaginae
 Familia: Silphidae Latreille, 1807 [Pg23 ... R] Pal/Rus
 Subfamilia: Nicrophorinae Kirby, 1837 [Pg2-3 ... R] Pal/Rus
 Subfamilia: Silphinae Latreille, 1802 [ ... R] Pal/Rus
 Familia: Scydmaenidae Leach, 1815 [K12 ... R] Pal/Rus
 =  Anisosphaeridae Temoesvary, 1882
 Subfamilia: Mastiginae Fleming, 1821 [ ... R] ?/?
 Subfamilia: Scydmaeninae Leach, 1815 [ ... R] Pal/Rus
 Familia: Ptiliidae Heer, 1843 [Pg23 ... R] Pal/Rus
 =  Cephaloplectidae Sharp, 1882
 =  Limulodidae Ganglbauer, 1899
 =  Trichopterigidae Erichson, 1845
 Subfamilia: Acrotrichinae Reitter, 1909 [ ... R] Pal/Rus
 Subfamilia: Cephaloplectinae Sharp, 1883 [ ... R] ?/?
 =  Limulodinae Ganglbauer, 1899
 Subfamilia: Nanosellinae Barber, 1924 [ ... R] ?/?
 Subfamilia: Ptiliinae Erichson, 1845 [ ... R] Pal/Rus
 Familia: Staphylinidae Latreille, 1802 [J12 ... R] Pal/Rus
 =  Brathinidae
 =  Dasyceridae Reitter, 1887
 =  Empelidae
 =  Micropeplidae Leach, 1815 ?(Heer, 1838)
 =  Oxyporidae Fleming, 1821
 =  Oxytelidae Fleming, 1821
 Subfamilia: Glypholomatinae Jeannel, 1962 [ ... ] -/-
 Subfamilia: Microsilphinae Crowson, 1950 [ ... R] -/-
 Subfamilia: Dasycerinae Reitter, 1887 [ ... R] Pal/Rus
 Subfamilia: Protopselaphinae Newton et Thayer, 1995 [ ... R] -/-
 Subfamilia: Pselaphinae Latreille, 1802 [Pg23 ... R] Pal/Rus
 =  Batrisinae Reitter, 1882 ?(Raffrai, 1890)
 =  Bythininae Raffrai, 1890
 =  Clavigerinae Leach, 1815
 =  Euplectinae
 =  Faroninae Reitter, 1882
 =  Goniacerinae Reitter, 1882
 =  Mayetiinae
 =  Pselaphinae Latreille, 1802
 Subfamilia: Trichophyinae C.G. Thomson, 1858 [ ... R] -/-
 Subfamilia: Phloeocharinae Erichson, 1839 [ ... R] Pal/Rus
 Subfamilia: Scaphidiinae Latreille, 1806 [Pg23 ... R] Pal/Rus
 Subfamilia: Piestinae Erichson, 1839 [ ... R] -/-
 Subfamilia: Aphaenostemminae Peyerimhoff, 1914 [ ... R] ?/?
 Subfamilia: Empelinae (Abdullah, 1969) Newton et Thayer, 1991 [ ... R] -/-
 Subfamilia: Omaliinae Macleay, 1825 [J13 ... R] Pal/Rus
 =  Brathinidae
 =  Aphaenostemminae
 Subfamilia: Proteininae Erichson, 1839 [ ... R] Pal/Rus
 =  Metopsiinae
 =  Pteroniinae
 Subfamilia: Micropeplinae Leach, 1815 [ ... R] Pal/Rus
 Subfamilia: Trilobitideinae Eichelbaum, 1909 [ ... ]
 Subfamilia: Pseudopsinae Ganglbauer, 1895 [ ... R] ?/?
 Subfamilia: Olisthaerinae C.G. Thomson, 1859 [J21 ... R] Pal/Rus
 Subfamilia: Solieriinae Newton et Thayer, 1991 [ ... R] -/-
 Subfamilia: Apateticinae Fauvel, 1895 [ ... R] ?/?
 Subfamilia: Trigonurinae Reiche, 1845 [J33 ... R] Pal/Rus
 Subfamilia: Osoriinae Erichson, 1839 [Pg23, BalJ ... R] ?/?
 Subfamilia: Neophoninae Fauvel, 1905 [ ... R] ?/?
 Subfamilia: Habrocerinae Mulsant et Rey, 1877 [ ... R] Pal/Rus
 Subfamilia: Aleocharinae Fleming, 1821 [ ... R] Pal/Rus
 =  Cyphinae
 =  Hypocyptinae Ganglbauer, 1895
 =  Mesoporinae
 =  Mimanommatinae Wasmann, 1912
 =  Pseudoperinthinae
 =  Pulicomorphinae Mann, 1921
 =  Pygosteninae Wasmann, 1902
 =  Termitodiscinae Wasmann, 1912
 =  Trichopseniinae Eichelbaum, 1909
 Subfamilia: Oxyporinae Fleming, 1821 [ ... R] Pal/Rus
 Subfamilia: Oxytelinae Fleming, 1821 [ ... R] Pal/Rus
 Subfamilia: Megalopsidiinae Leng, 1920 [ ... R] -/-
 Subfamilia: Steninae Macleay, 1825 [ ... R] Pal/Rus
 Subfamilia: Euaesthetinae C.G. Thomson, 1859 [ ... R] ?/?
 Subfamilia: Leptothyphlinae Fauvel, 1874 [ ... R] ?/?
 Subfamilia: Trichophyinae C.G. Thomson, 1859 [ ... R] -/-
 Subfamilia: Tachyporinae MacLeay, 1825 [J31, KarT ... R] Pal/Rus
 Subfamilia: Paederinae Ganglbauer, 1895 [Pg22, GreR ... R] Pal/Rus
 Subfamilia: Staphylininae Latreille, 1802 [ ... R] Pal/Rus
 =  Amblyopininae
 =  Diochinae
 =  Platyprosopinae
 =  Quedinae Ganglbauer, 1895
 =  Xantholininae Ganglbauer, 1895
 =  Xanthoperiginae
 Superfamilia: Scarabaeoidea Latreille, 1802 [ ... ] Pal/Rus
 =  Lamellicornia
 Familia: Glaresidae Semenov-Tian-Shanskii et Medvedev, 1932 [ ... R] Pal/?
 Familia: Passalidae Leach, 1815 [ ... R] -/-
 Subfamilia: Aulacocyclinae Kaup, 1868 [Pg33 ... R] -/-
 Subfamilia: Passalinae Leach, 1815 [Post, N12, Sha ... R] -/-
 Familia: Lucanidae Latreille, 1804 [K2t ... R] Pal/Rus
 Subfamilia: Aesalinae Latreille, 1804 [ ... R] Pal/Rus
 Subfamilia: Nicaginae LeConte, 1861 [ ... R] Pal/?
 Subfamilia: Syndesinae MacLeay, 1819 [Pg31 ... R] Pal/Rus
 =  Sinodendrinae
 Subfamilia: Lampriminae MacLeay, 1819 [Pg23, BalJ ... R] -/-
 Subfamilia: Penichrolucaninae Arrow, 1950 [ ... R] -/-
 Subfamilia: Lucaninae Latreille, 1804 [Pg23 ... R] Pal/Rus
 =  Chiasognathinae
 =  Dorcinae
 =  Figulinae
 =  Odontolabinae
 =  Platycerinae
 Familia: Trogidae Macleay, 1819 [ ... ] Pal/Rus
 Familia: Pleocomidae LeConte, 1861 [ ... R] -/-
 Familia: Diphyllostomatidae Holloway, 1972 [ ... R] -/-
 Familia: Geotrupidae Latreille, 1802 [K12 ... R] Pal/Rus
 Subfamilia: Bolboceratinae Mulsant, 1842 [K12 ... R] ?/?
 =  Athyreinae Howden et Martinetz, 1963
 =  Bolbocerinae
 Subfamilia: Geotrupinae Latreille, 1802 [ ... R] Pal/Rus
 =  Taurocerastinae
 Subfamilia: Lethrinae Mulsdant et Rey, 1871 [ ... R] Pal/Rus
 Familia: Ochodaeidae Mulsant et Rey, 1871 [K1ap ... R] Pal/Rus
 Subfamilia: Ochodaeinae Mulsant et Rey, 1871 [ ... R] Pal/Rus
 Subfamilia: Chaetocanthinae Scholtz in Scholtz et al., 1988 [ ... R] -/-
 Familia: Belohinidae Paulian, 1959 [ ... R] ?/?
 Familia: Ceratocanthidae Cartwright et Gordon, 1971 [ ... R] -/-
 =  Acanthoceridae
 Familia: Glaphyridae Macleay, 1819 [ ... R] ?/?
 =  Lichniinae
 Familia: Scarabaeidae Latreille, 1802 [J2 ... R] Pal/Rus
 =  Aclopidae
 =  Cetoniidae
 Subfamilia: Lithoscarabaeinae Nikolajev, 1992 [K12, Bais ... R] Pal/?
 Subfamilia: Hybosorinae Erichson, 1847 [K12 ... R] ?/?
 Subfamilia: Aclopinae Milne-Edwards, 1850 [J3ox ... R] -/-
 Subfamilia: Scarabaeinae Latreille, 1802 [Pg1d ... R] Pal/Rus
 Subfamilia: Coprinae [K2tu ... R] Pal/Rus
 Subfamilia: Aphodiinae Leach, 1815 [K12 ... R] Pal/Rus
 =  Aegaliinae
 =  Aulonocneminae
 =  Chironinae
 =  Silluviinae
 =  Termithtroginae
 Subfamilia: Pachypodinae Erichson, 1840 [ ... R] Pal/Rus
 Subfamilia: Oncerinae [ ... R] ?/?
 Subfamilia: Orphninae Erichson, 1847 [ ... R] ?/?
 Subfamilia: Allidostomatinae Arrow, 1940 [ ... R] -/-
 =  Allidiostominae
 =  Idiostominae
 Subfamilia: Dynamopodinae Arrow, 1911 [ ... R] Pal/?
 =  Dynamopinae
 Subfamilia: Melolonthinae MacLeay, 1819 [Pg21, Wilc ... R] Pal/Rus
 =  Syntellopodinae
 Subfamilia: Sericinae [K12 ... R] Pal/Rus
 Subfamilia: Hopliinae [Pg31, Flor ... R] Pal/Rus
 Subfamilia: Euchirinae Hope, 1840 [N12, Tott ... R] Pal/?
 Subfamilia: Phenomeridinae Erichson, 1847 [ ... R] ?/?
 =  Phenomerinae
 Subfamilia: Cetoniinae Leach, 1815 [N12 ... R] Pal/Rus
 =  Trichinae
 =  Valginae
 Subfamilia: Dynastinae MacLeay, 1819 [Pg31 ... R] ?/?
 Subfamilia: Rutelinae MacLeay, 1819 [Pg22, Mess ... R] Pal/Rus
 Infraordo: Elateriformia [ ... ] Pal/Rus
 =  Dascilliformia
 =  Eucinetiformia
 Superfamilia: Dascilloidea Gue'rin-Me'neville, 1843 [ ... ] Pal/Rus
 Familia: Dascilidae Gue'rin-Me'neville, 1843 [K2cm ... R] Pal/Rus
 =  Karumiidae Escalera, 1913
 Subfamilia: Dascillinae Gue'rin-Me'neville, 1843 [K2cm ... R] Pal/Rus
 =  Atopinae
 =  Cinnabariinae
 Subfamilia: Karumiinae Escalera, 1913 [ ... R] Pal/?
 =  Zarudniolidae
 Familia: Rhipiceridae Latreille, 1834 [ ... R] Pal/Rus
 =  Sandalidae
 Superfamilia: Scirtoidea Fleming, 1821 [ ... ] Pal/Rus
 =  Clamboidea Fischer, 1821
 =  Eucinetoidea Lacordaire, 1857
 Familia: Decliniidae Nikit., Lawr., Kirejt. et Gratsh., 1994 [ ... R] Pal/Rus
 Familia: Eucinetidae Lacordaire, 1857 [J3ox ... R] Pal/Rus
 =  Cryptomeridae
 Familia: Clambidae Fischer, 1821 [Pg23 ... R] Pal/Rus
 =  Calyptomeridae
 Subfamilia: Calyptomerinae Crowson, 1955 [ ... R] ?/?
 Subfamilia: Acalyptomerinae [ ... R] ?/?
 Subfamilia: Clambinae Fischer, 1821 [ ... R] Pal/Rus
 Familia: Scirtidae Fleming, 1821 [Pg23 ... R] Pal/Rus
 =  Cyphonidae Stephens, 1829
 =  Elodidae Shuckard, 1840
 =  Helodidae Agassiz, 1846
 Superfamilia: Buprestoidea [ ... ] Pal/Rus
 Familia: Buprestidae Leach, 1815 [J2bt ... R] Pal/Rus
 =  Electrapatidae Iablokoff-Khnzorian, 1962
 Subfamilia: Schizopodinae Leconte, 1861 [ ... P] -/-
 Subfamilia: Julodinae Lacordaire, 1857 [ ... P] Pal/?
 Subfamilia: Parathyreinae Alexeev, 1993 [J2bt ... K1]
 Subfamilia: Chalcophorinae Lacordaire, 1857 [Pg22 ... R] Pal/Rus
 Subfamilia: Buprestinae Leach, 1815 [Pg13, Mena ... R] Pal/Rus
 =  Chrysobothrinae
 Subfamilia: Mastogeniinae Leconte et Horn, 1883 [Pg23 ... R]
 Subfamilia: Sphenopterinae Lacordaire, 1857 [Pg22, Mess ... R] Pal/Rus
 Subfamilia: Agrilinae Laporte, 1835 [Pg ... R] Pal/Rus
 =  Galbellinae
 Subfamilia: Polycestinae Lacordaire, 1857 [ ... P] ?/?
 Subfamilia: Acmaeoderinae [Pg13, Mena ... R] Pal/Rus
 Subfamilia: Sternocerinae [ ... ]
 Subfamilia: Thrincopyginae LeConte, 1861 [ ... R] ?/?
 Subfamilia: Cylindromorphinae Portevin, 1931 [ ... R] Pal/-
 Subfamilia: Trachyinae Laporte, 1835 [ ... R] Pal/Rus
 Superfamilia: Byrrhoidea [ ... ] Pal/Rus
 =  Dryopoidea
 =  Psephenoidea
 Familia: Byrrhidae Latreille, 1806 [J11 ... R] Pal/Rus
 =  Syncaliptidae Mulsant et Rey, 1869
 Subfamilia: Byrrinae Latreille, 1806 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Amphicyrtinae Leconte, 1861 [Pg31, Flor ... R] ?/?
 Subfamilia: Syncaliptinae Mulsant et Rey, 1869 [Pg23, BalJ ... R] ?/?
 Familia: Elmidae Curtis, 1830 [Pg22 ... R] Pal/Rus
 =  Elmididae
 =  Elminthidae
 =  Helminthidae
 =  Linmiidae
 Subfamilia: Larainae LeConte, 1861 [Pg22, Geis ... R] ?/?
 =  Larinae
 Subfamilia: Elminae Curtis, 1830 [Pg23 ... R] Pal/Rus
 Familia: Dryopidae Billberg, 1820 [Pg31 ... R] Pal/Rus
 =  Chiloeidae
 =  Parnidae
 Familia: Lutrochidae Kasap et Crowson, 1975 [Pg31 ... R] -/-
 Familia: Limnichidae Erichson, 1846 [Pg23 ... R] Pal/Rus
 Subfamilia: Hyphalinae Britton, 1971 [ ... ]
 Subfamilia: Limnichinae Erichson, 1846 [Pg23 ... R] Pal/Rus
 Subfamilia: Cephalobyrrhinae Champion, 1925 [ ... R] ?/?
 Subfamilia: Thaumastodinae Champion, 1924 [ ... R] ?/?
 Familia: Heteroceridae MacLeay, 1825 [K11 ... R] Pal/Rus
 Subfamilia: Elythomerinae Pacheco, 1964 [ ... R] ?/?
 Subfamilia: Heterocerinae MacLeay, 1825 [K11 ... R] Pal/Rus
 Familia: Psephenidae Lacordaire, 1854 [Pg22 ... R] Pal/Rus
 =  Eubriidae
 Familia: Psephenoididae [ ... ]
 Subfamilia: Eubriinae Lacordaire, 1857 [ ... R] Pal/Rus
 Subfamilia: Psephenoidinae Hinton, 1939 [ ... R] ?/?
 Subfamilia: Eubrianacinae Jacobson, 1913 [ ... R] ?/?
 =  Placonychinae Horn, 1880
 Subfamilia: Psepheninae Lacordaire, 1854 [Pg31, Flor ... R] Pal/Rus
 Familia: Cneoglossidae Champion, 1897 [ ... R] -/-
 Familia: Ptylodactylidae Laporte, 1836 [Pg23 ... R] Pal/Rus
 Subfamilia: Anchytarsinae Champion, 1897 [ ... R] Pal/-
 Subfamilia: Cladotominae Pic, 1914 [ ... R] Pal/-
 Subfamilia: Aploglossinae Champion, 1897 [ ... R] -/-
 Subfamilia: Araeopidiinae Lawrence, 1991 [ ... R] -/-
 Subfamilia: Ptylodactylinae Laporte, 1836 [Pg23 ... R] Pal/Rus
 Familia: Chelonariidae Blanchard, 1845 [Pg31 ... R] -/-
 Familia: Eulichadidae Crowson, 1973 [ ... R] Pal/-
 =  Lichadidae
 Familia: Callirhipidae Emden, 1924 [ ... R] -/-
 Superfamilia: Cebrionoidea [ ... ] Pal/
 =  Elateroidea
 Familia: Artematopodidae Lacordaire, 1857 [Pg23, BalJ ... R] Pal/Rus
 =  Artematopidae
 =  Eurypogonidae
 Subfamilia: Electribiinae Crowson, 1973 [Pg23, BalJ ... R] Pal/?
 Subfamilia: Artematopinae Lacordaire, 1857 [Pg23, BalJ ... R] Pal/Rus
 Familia: Brachypsectridae LeConte et Horn, 1883 [Pg33 ... R] -/-
 Familia: Rhinorhipidae Lawrence, 1987 [ ... R] -/-
+Familia: Praelateriidae Dolin, 1973 [J1he ... J1he] Pal/Rus
 Familia: Elateridae Leach, 1815 [J1he ... R] Pal/Rus
 =  Dicronychidae
 =  Lissomidae
 Subfamilia: Tetralobinae Laporte, 1840 [ ... R] -/-
 Subfamilia: Thylacosterninae Fleutiaux, 1920 [ ... R] ?/?
 =  Balginae
 =  Soleniscinae
 Subfamilia: Lissominae Laporte, 1835 [Pg1dn ... R] Pal/Rus
 Subfamilia: Oxynopterinae Cande`ze, 1857 [N1aq ... R] ?/?
 Subfamilia: Semiotinae Jacobson, 1913 [Pg1 ... R] ?/?
 Subfamilia: Pityobiinae Hyslop, 1917 [ ... R] -/-
+Subfamilia: Protagrypninae Dolin, 1973 [J1he ... J3ox] Pal/?
 Subfamilia: Agripninae Cande`ze, 1857 [J3ox ... R] Pal/Rus
 =  Cavicoxumidae
 =  Pangauridae Gistel, 1856
 =  Adelocerinae
 =  Campyloxeninae
 =  Hemirhipinae
 =  Pyrophorinae
 Subfamilia: Pyrophorinae Cande`ze, 1863 [Pg22 ... R]
 Subfamilia: Melanactinae Cande`ze, 1857 [ ... R]
 Subfamilia: Hypnoidinae Schwarz, 1906 [Pg23 ... R] Pal/Rus
 =?  Hypolithinae
 Subfamilia: Denticollinae Stein et Weise, 1877 [J3ox, KarT ... R] Pal/Rus
 =  Athoinae
 =  Campylinae
 =  Crepidomeninae
 =  Melanactinae
 Subfamilia: Diminae Cande`ze, 1859 [ ... R] ?/?
 Subfamilia: Elaterinae Leach, 1815 [Pg2 ... R] Pal/Rus
 =  Abrastinae
 Subfamilia: Hemiopinae Feutiaux, 1941 [ ... R] ?/?
 =  Hemiopsinae
 Subfamilia: Pleonominae Semenov-Tian-Shanskii et Piatakov, 1936 [ ... R]
 Subfamilia: Physodactylinae Lacordaire, 1857 [ ... R] ?/?
 Subfamilia: Aplastinae Stibick, 1979 [ ... R] ?/?
 Subfamilia: Oestodinae Hyslop, 1917 [K12 ... R] ?/?
 Subfamilia: Melanotinae Cande`ze, 1859 [ ... R] Pal/Rus
 Subfamilia: Negastriinae Nakane et Kishii, 1956 [J3ox ... R] Pal/Rus
 Subfamilia: Cardiophorinae Cande`ze, 1860 [J3ox ... R] Pal/Rus
 Subfamilia: Eudicronychinae Girard, 1971 [ ... R] ?/?
 =  Dicronychinae Schwartz, 1897
 Subfamilia: Anischiinae Feutiaux, 1936 [ ... R] ?/?
 Subfamilia: Suprotelaterinae Feutiaux, 1920 [ ... R] ?/?
 Familia: Eucnemidae Eschscholtz, 1829 [Pg23 ... R] Pal/Rus
 =  Melasidae Fleming, 1821
 =  Perothopidae Lacordaire, 1857
 =  Phylloceridae Reitter, 1905
 Subfamilia: Perothopinae Lacordaire, 1857 [ ... R] -/-
 Subfamilia: Phyllocerinae Reitter, 1905 [ ... R] Pal/-
 Subfamilia: Pseudomeninae Muona, 1993 [ ... R] -/-
 Subfamilia: Palaeoxeninae Muona, 1993 [ ... R] -/-
 Subfamilia: Phlegoninae Muona, 1993 [ ... R] -/-
 Subfamilia: Melasinae Fleming, 1821 [Pg31, Flor ... R] Pal/Rus
 Subfamilia: Eucneminae Eschscholtz, 1829 [ ... R] Pal/Rus
 Subfamilia: Macraulacinae Fleutiaux, 1923 [Pg23, BalJ ... R] ?/?
 Familia: Cebrionidae Latreille, 1802 [ ... R] Pal/-
 =  Aplastinae
 =  Cebriognathinae
 =  Plastocerinae
 Familia: Cerophytidae Latreille, 1834 [K12 ... R] Pal/Rus
 Familia: Throscidae Laporte, 1840 [K12 ... R] Pal/Rus
 =  Trixagidae
 Familia: Plastoceridae Crowson, 1972 [ ... R] Pal/-
 Superfamilia: Cantharoidea [ ... ] Pal/Rus
 Familia: Omalisidae Lacordaire, 1857 [ ... R] ?/?
 =  Homalisidae
 =  Omalysidae
 Familia: Berendtimiridae Winkler, 1987 [Pg23, BalJ ... R] ?/?
 Familia: Lycidae Laporte, 1836 [Pg23 ... R] Pal/Rus
 Subfamilia: Lycinae Laporte, 1836 [ ... R] Pal/Rus
 Subfamilia: Leptolycinae Leng et Mutchler, 1922 [ ... R] ?/?
 =  Duliticolinae Mjoeberg, 1925
 Subfamilia: Ateliinae Kleine, 1928 [ ... R] ?/?
 Subfamilia: Metriorrhynchinae Kleine, 1926 [ ... R] ?/?
 Subfamilia: Erotinae Leconte, 1881 [Pg23 ... R] Pal/Rus
 Subfamilia: Calochrominae Lacordaire, 1857 [ ... R] Pal/Rus
 Familia: Drilidae Blanchard, 1845 [ ... R] Pal/Rus
 Familia: Telegeusidae Leng, 1920 [ ... R] -/-
 Familia: Lampyridae Latreille, 1817 [Pg23 ... R] Pal/Rus
 Subfamilia: Pterotinae LeConte, 1861 [ ... R] -/-
 Subfamilia: Cyphonocerinae Crowson, 1972 [ ... R] ?/-
 Subfamilia: Ototretadrilinae Crowson, 1972 [ ... R] ?/-
 Subfamilia: Ototretinae McDermott, 1964 [ ... R] -/-
 Subfamilia: Amydetinae Olivier, 1907 [ ... R] ?/?
 Subfamilia: Lampyrinae Latreille, 1817 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Luciolinae Lacordaire, 1857 [Pg23 ... R] Pal/Rus
 Subfamilia: Photurinae Lacordaire, 1857 [ ... R] ?/?
 Familia: Omethidae LeConte, 1861 [ ... R] Pal/-
 Subfamilia: Driloniinae Crowson, 1972 [ ... R] -/-
 Subfamilia: Matheteinae LeConte, 1861 [ ... R] -/-
 Subfamilia: Omethinae LeConte, 1861 [ ... R] Pal/-
 Familia: Phengodidae LeConte, 1861 [ ... R] -/-
 Subfamilia: Rhagophthalminae Olivier, 1907 [ ... R] -/-
 Familia: Phengodidae LeConte, 1861 [ ... R] -/-
 Familia: Cantharidae Latreille, 1802 [Pg23 ... R] Pal/Rus
 =  Chaulignathidae
 =  Telephoridae
 Subfamilia: Cantharinae Latreille, 1802 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Dysmorphocerinae Brancucci, 1980 [ ... R] ?/?
 Subfamilia: Malthininae Kiesenwetter, 1852 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Chaulignathinae LeConte, 1861 [Pg31, Flor ... R] ?/?
 Subfamilia: Silinae Mulsant, 1862 [ ... R] Pal/Rus
 Familia: Podabrocephalidae Pic, 1930 [ ... R] -/-
 Infraordo: Cucujiformia [ ... ] Pal/Rus
 =  Chrysomelomorpha
 Superfamilia: Derodontoidea LeConte, 1861 [ ... R] Pal/Rus
 Familia: Derodontidae LeConte, 1861 [ ... R] Pal/Rus
 =  Laricobiidae Mulsant et Rey, 1863-1864
 =  Peltasticidae LeConte, 1861
 Subfamilia: Peltasticinae LeConte, 1861 [ ... R] Pal/Rus
 Subfamilia: Derodontinae LeConte, 1861 [ ... R] Pal/Rus
 Subfamilia: Laricobiinae Mulsant et Rey, 1863-1864 [ ... R] Pal/Rus
 Superfamilia: Bostrichoidea [K12 ... R] Pal/Rus
 =  Dermestoidea
 Familia: Nosodendridae Erichson, 1846 [Pg22 ... R] Pal/Rus
 Familia: Dermestidae Latreille, 1807 [K12 ... R] Pal/Rus
 =  Thorictidae
 =  Thylodriinae
 Subfamilia: Dermestinae Latreille, 1807 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Attageninae Laporte, 1840 [Pg23 ... R] Pal/Rus
 Subfamilia: Megatominae Leach, 1815 [Pg23 ... R] Pal/Rus
 Subfamilia: Orphilinae Leconte, 1861 [Pg31 ... R] Pal/Rus
 Subfamilia: Trinodinae Casey, 1900 [Pg23 ... R] Pal/Rus
 Subfamilia: Egidyellinae ???? [ ... R] Pal/-
 Subfamilia: Marioutinae Jacobson, 1913 [ ... R] Pal/-
 Subfamilia: Thorictinae Agassiz, 1846 [ ... R] Pal/?
 Subfamilia: Thylodriinae Semenov-Tian-Shanskii, 1913 [ ... R] Pal/Rus
 Familia: Endecotomidae LeConte, 1861 [ ... R] Pal/Rus
 Familia: Bostrichidae Latreille, 1802 [Pg21, LonC ... R] Pal/Rus
 =  Bostrychidae
 =  Lyctidae
 =  Psoidae
 Subfamilia: Dysidinae Lesne, 1921 [ ... R] ?/?
 =  Apoleoninae
 Subfamilia: Polycaininae Lesne, 1896 [ ... R] ?/?
 Subfamilia: Psoinae Blanchard, 1851 [ ... R] Pal/Rus
 Subfamilia: Dinoderinae C.G. Thomson, 1863 [Pg31 ... R] ?/?
 Subfamilia: Lyctinae Billberg, 1820 [Pg23 ... R] Pal/Rus
 Subfamilia: Euderinae Lesne, 1834 [ ... R] -/-
 Subfamilia: Bostrichinae Latreille, 1802 [Pg23 ... R] Pal/Rus
 Familia: Anobiidae Stephens, 1830 [K12 ... R] Pal/Rus
 =  Ectrephidae
 =  Gnostidae
 =  Ptinidae Latreille, 1802
 Subfamilia: Eucradinae LeConte, 1861 [Pg23 ... R] Pal/Rus
 =  Hedobiinae Mulsant et Rey, 1868
 Subfamilia: Ptininae Latreille, 1802 [Pg23 ... R] Pal/Rus
 =  Gibbiinae Mulsant et Rey, 1868
 Subfamilia: Ptilininae Shuckard, 1840 [Pg23 ... R] Pal/Rus
 Subfamilia: Dryophilinae LeConte, 1861 [ ... R] Pal/Rus
 Subfamilia: Alvarenganellinae Viana et Martinez, 1971 [ ... R] -/-
 Subfamilia: Ernobiinae Pic, 1912 [ ... R] Pal/Rus
 =  Cerocosminae Pic, 1912
 =  Cosmocerinae Solier, 1849
 Subfamilia: Xyletininae Gistel, 1856 [Pg23 ... R] Pal/Rus
 Subfamilia: Dorcatominae C.G. Thomson, 1859 [Pg23 ... R] Pal/Rus
 =  Dorkatominae
 Subfamilia: Mesocoelopodinae Mulsant et Rey, 1864 [Pg23 ... R] Pal/Rus
 =  Mesothinae Portevin, 1931
 =  Tricoryninae White, 1971
 Familia: Jacobsoniidae Heller, 1926 [ ... ]
 =  Sarothriidae Crowson, 1955
 =  Derolathrinae Sen Gupta, 1979
 Superfamilia: Lymexyloidea Fleming, 1921 [ ... ] Pal/Rus
 Familia: Lymexylidae Fleming, 1921 [Pg23 ... R] Pal/Rus
 =  Atractoceridae Laporte, 1840
 =  Lymexylonidae
 Subfamilia: Lymexylinae Fleming, 1921 [Pg23 ... R] Pal/Rus
 Subfamilia: Hylecoetinae Gistel, 1856 [N13 ... R] Pal/Rus
 Subfamilia: Melittommatinae Wheeler, 1986 [ ... R] -/-
 Superfamilia: Cleroidea Latreille, 1802 [J1to ... R] Pal/Rus
 Familia: Phloiophilidae Kisenwetter, 1863 [ ... R] Pal/?
 =  Phloeophilidae Kisenwetter, 1863
 Familia: Peltidae Kirby, 1837 [J1to ... R] Pal/Rus
 =  Larinotidae Slipinski, 1992
 =  Ostomidae Reitter, 1882
 Familia: Lophocateridae [ ... ] Pal/Rus
+Subfamilia: Meligethiellinae Kirejtshuk et Ponomarenko, 1990 [J31 ... ???] Pal/?
 Subfamilia: Protopeltinae Crowson, 1966 [ ... R] -/-
 Subfamilia: Larinotinae Slipinski, 1992 [ ... R] -/-
 Subfamilia: Rentoniinae Crowson, 1966 [ ... R] -/-
 Subfamilia: Decamerinae Crowson, 1964 [ ... R] -/-
 Subfamilia: Peltinae Latreille, 1807 [Pg23 ... R] Pal/Rus
 =  Ostominae
 Subfamilia: Lophocaterinae Crowson, 1964 [Pg23 ... R] Pal/Rus
 Familia: Trogossitidae Latreille, 1802 [ ... R] Pal/Rus
 =  Nemosomatidae Leach, 1815
 =  Nemosomidae
 =  Temnochilidae
 =  Tenebroididae
 Subfamilia: Gymnochilinae Lacordaire, 1854 [Pg23 ... R] Pal/Rus
 =  Calytinae Reitter, 1922
 Subfamilia: Egoliinae Lacordaire, 1854 [ ... R] ?/?
 Subfamilia: Trogossitinae Latreille, 1802 [Pg1d ... R] Pal/Rus
 =  Nemosominae
 =  Temnochilinae
 =  Tenebroidinae
 Familia: Chaetosomatidae Crowson, 1955 [ ... R] Pal/Rus
 Familia: Cleridae Latreille, 1802 [Pg13 ... R] Pal/Rus
 =  Corynetidae Laporte, 1836
 =  Korynetidae
 =  Thanerocleridae Chapin, 1924
 Subfamilia: Thaneroclerinae Chapin, 1924 [ ... R] ?/?
 Subfamilia: Tillinae Leach, 1815 [Pg23 ... R] Pal/Rus
 Subfamilia: Hydnocerinae Spinola, 1844 [Pg23 ... R] ?/?
 =  Phyllobaeninae Lacordaire, 1857
 Subfamilia: Clerinae Latreille, 1802 [Pg13 ... R] Pal/Rus
 =  Cleropiestinae Winkler, 1980
 =  Dieropsinae Winkler, 1964
 =  Priocerinae
 Subfamilia: Epiphleinae Kuwert, 1893 [ ... R] -/-
 Subfamilia: Tarsosteninae Jacquelin du Val, 1860 [Pg23 ... R] -/-
 Subfamilia: Korynetinae Laporte, 1836 [Pg23 ... R] Pal/Rus
 =  Corynetinae
 Familia: Acanthocnemidae Crowson, 1964 [ ... R] -/-
 Familia: Phycosecidae Crowson, 1952 [ ... R] -/-
 Familia: Prionoceridae Lacordaire, 1857 [ ... R] Pal/?
 Familia: Melyridae Leach, 1815 [ ... ] Pal/Rus
 =  Attalomenidae Majer, 1994
 =  Carphuridae
 =  Dasytidae Laporte, 1840
 =  Malachiidae Fleming, 1821
 =  Rhadalidae LeConte, 1861
 Subfamilia: Melyrinae Leach, 1815 [Pg23 ... R] Pal/Rus
 =  Attalomenidae Majer, 1994
 Subfamilia: Rhadalinae LeConte, 1861 [Pg23 ... R] Pal/?
 =  Aplocneminae
 =  Haplocneminae
 Subfamilia: Gietellinae Constantin de Minier, 1987 [ ... R] Pal/-
 Subfamilia: Dasytinae Laporte, 1840 [Pg23 ... R] Pal/Rus
 Subfamilia: Malachiinae Fleming, 1821 [ ... R] Pal/Rus
 Familia: Mauroniscidae Majer, 1994 [ ... R] -/-
 Superfamilia: Cucujoidea [ ... ] Pal/Rus
 =  Clavicornia
+Familia: Parandrexidae Kirejtshuk, 1993 [J21 ... J3ox] Pal/-
 Familia: Protocucujidae Crowson, 1954 [ ... R] -/-
 Familia: Sphindidae Jacquelin du Val, 1858 [Pg23 ... R] Pal/Rus
 =  Aspidiphoridae Kiesenwetter, 1877
 Subfamilia: Protosphindinae Sen Gupta et Crowson, 1979 [ ... R] -/-
 Subfamilia: Odontosphindinae Sen Gupta et Crowson, 1979 [ ... R] Pal/-
 Subfamilia: Sphindiphorinae Sen Gupta et Crowson, 1979 [ ... R] -/-
 Subfamilia: Sphindinae Jacquelin du Val, 1858 [Pg23 ... R] Pal/Rus
 =  Aspidiphoridae Kiesenwetter, 1877
 =  Eurysphindinae
 Familia: Helotidae Reitter, 1876/Chapuis, 1876 [ ... R] Pal/Rus
 Familia: Nitidulidae Latreille, 1802 [K12 ... R] Pal/Rus
 =  Carpophilidae Murray, 1864
 =  Cybocephalidae Jacquelin du Val, 1858
 =  Ipidae Lacordaire, 1854
 =  Mystropidae Murray, 1864
 =  Strongylinae Murray, 1864
 Subfamilia: Maynipeplinae Kirejtshuk, 1998 [ ... R] -/-
 Subfamilia: Calonecrinae Kirejtshuk, 1982 [ ... R] -/-
 Subfamilia: Epuraeinae Kirejtshuk, 1986 [ ... R] Pal/Rus
 Subfamilia: Carpophilinae Erichson, 1842 [ ... R] Pal/Rus
 Subfamilia: Amphicrossinae Kirejtshuk, 1986 [ ... R] Pal/Rus
 Subfamilia: Meligethinae C.G. Thomson, 1859 [Pg23 ... R] Pal/Rus
 Subfamilia: Nitidulinae Latreille, 1802 [K2cm ... R] Pal/Rus
 =  Strongylinae Erichson, 1843
 Subfamilia: Cillaeinae Kirejtshuk et Audisio, 1986 [Pg31 ... R] Pal/Rus
 Subfamilia: Cryptarchinae C.G. Thomson, 1859 [Pg23 ... R] Pal/Rus
 =  Ipinae Erichson, 1843
 =  Ipsinae Sharp, 1891
 =  Pityophaginae Fauconnet, 1894
 Subfamilia: Cybocephalinae Jacquelin du Val, 1858 [Pg23 ... R] Pal/Rus
 Familia: Boganiidae Sen Gupta et Crowson, 1966 [ ... R] -/-
 Subfamilia: Paracucujinae Endroedy-Younga et Crowson, 1986 [ ... R] -/-
 Subfamilia: Boganiinae Sen Gupta et Crowson, 1966 [ ... R] -/-
 Familia: Monotomidae Laporte, 1840 [Pg23 ... R] Pal/Rus
 =  Rhizophagidae Redtenbacher, 1845
 Subfamilia: Rhizophaginae Redtenbacher, 1845 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Monotominae [ ... ] Pal/Rus
 =  Lenacinae
 =  Thioninae
 Familia: Kateretidae Erichson, 1844 [Pg31 ... R] Pal/Rus
 =  Brachypteridae Erichson, 1845
 Familia: Smicripidae Horn, 1879 [ ... R] -/-
 Familia: Phloeostichidae Reitter, 1911 [ ... R] Pal/Rus
 Subfamilia: Phloestichinae Reitter, 1911 [ ... R] Pal/Rus
 Subfamilia: Hymaeinae Sen Gupta et Crowson, 1966 [ ... R] -/-
 Subfamilia: Agapythinae Sen Gupta et Crowson, 1966 [ ... R] -/-
 Subfamilia: Myraboliinae Lawrence et Britton, 1991 [ ... R] -/-
 Subfamilia: Priasilphinae Crowson, 1973 [ ... R] -/-
 Subfamilia: Tasmosalpinginae Lawrence et Britton, 1991 [ ... R] -/-
 Familia: Silvanidae Kirby, 1837 [Pg23 ... R] Pal/Rus
 Subfamilia: Brontinae Erichson, 1845/Blanchard, 1845 [ ... R] Pal/Rus
 =  Cryptamorphinae,
 =  Hyliotinae Reitter, 1879
 =  Psammoecinae
 =  Uleiotinae Gistel, 1856
 Subfamilia: Silvaninae Kirby, 1837 [Pg23 ... R] Pal/Rus
 Familia: Passandridae Erichson, 1845/Blanchard, 1845 [Pg23 ... R] Pal/?
 =  Catogenidae
 Familia: Cucujidae [ ... R] Pal/Rus
 Familia: Laemophloeidae Ganglbauer, 1899 [Pg22 ... R] Pal/Rus
 Familia: Propalticidae Crowson, 1952 [N2 ... R] -/-
 Familia: Phalacridae Leach, 1815 [Pg23 ... R] Pal/Rus
 =  Phaenocephalidae Matthews, 1899
 Subfamilia: Phaenocephalinae Matthews, 1899 [ ... R] Pal/-
 Subfamilia: Phalacrinae Leach, 1815 [Pg23 ... R] Pal/Rus
 Familia: Hobartiidae Sen Gupta et Crowson, 1966 [ ... ] -/-
 Familia: Cavognathidae Sen Gupta et Crowson, 1966 [ ... ] -/-
 Familia: Cryptophagidae Kirby, 1837 [K2sn ... R] Pal/Rus
 =  Catopochrotidae
 =  Hypocopridae Reitter, 1879
 Subfamilia: Hypocoprinae Reitter, 1879 [ ... R] Pal/Rus
 Subfamilia: Cryptophaginae Kirby, 1837 [Pg13 ... R] Pal/Rus
 =  Telmatophilinae
 Subfamilia: Atomariinae LeConte, 1861 [K2sn ... R] Pal/Rus
 =  Ephisteminae
 Familia: Lamingtoniidae Sen Gupta et Crowson, 1966 [ ... R] -/-
 Familia: Languriidae Crotch, 1873 [K1ap ... R] Pal/Rus
 Familia: Cryptophilidae Casey, 1900 [ ... R] Pal/Rus
 Subfamilia: Xenoscelinae Ganglbauer, 1899 [K1ap, Leba ... R] ?/?
 =  Loberinae
 =  Pharaxonotinae
 Subfamilia: Setariolinae Crowson, 1952 [ ... R] Pal/?
 =  Setariinae
 Subfamilia: Languriinae Crotch, 1873 [ ... R] Pal/Rus
 =  Cladoxeninae
 Subfamilia: Cryptophilinae Casey, 1900 [Pg23 ... R] Pal/Rus
 Subfamilia: Toraminae Sen Gupta, 1967 [ ... R] ?/?
 Familia: Erotylidae Latreille, 1802 [Pg23 ... R] Pal/Rus
 Familia: Dacnidae Gistel, 1856 [ ... ] Pal/Rus
 Subfamilia: Dacninae Gistel, 1856 [ ... R] Pal/Rus
 Subfamilia: Megalodacninae Sen Gupta, 1969 [ ... R] Pal/Rus
 Subfamilia: Encaustinae Crotch, 1876 [ ... R] ?/?
 Subfamilia: Tritominae Curtis, 1834 [Pg31 ... R] Pal/Rus
 =  Triplacinae Imhoff, 1856
 Subfamilia: Erotylinae Latreille, 1802 [ ... R] Pal/Rus
 Familia: Byturidae Jacquelin du Val, 1858 [K11 ... R] Pal/Rus
 Subfamilia: Platydascilinae Pic, 1914 [ ... R] ?/?
 Subfamilia: Byturinae Jacquelin du Val, 1858 [K11 ... R] Pal/Rus
 Familia: Biphyllidae LeConte, 1861 [ ... R] Pal/Rus
 =  Diphillidae
 Familia: Bothrideridae Erichson, 1845 [ ... R] Pal/Rus
 =  Anommatidae Ganglbauer, 1899
 Subfamilia: Teredinae Seidlitz, 1888 [ ... R] Pal/Rus
 Subfamilia: Xylariophilinae Pal et Lawrence, 1896 [ ... R] ?/?
 Subfamilia: Anommatinae Ganglbauer, 1899 [ ... R] ?/?
 Subfamilia: Bothriderinae Erichson, 1845 [Pg23 ... R] Pal/Rus
 =  Deretaphrinae
 Familia: Cerylonidae Billberg, 1820 [Pg23 ... R] Pal/Rus
 =  Aculagnathidae
 =  Cerylidae Billberg, 1820
 =  Dolosidae
 =  Euxestidae Grouvelle, 1908
 =  Murmidiidae Jacquelin du Val, 1858 (?)
 Subfamilia: Euxestinae Grouvelle, 1908 [ ... R] ?/?
 =  Metaceryloninae
 Subfamilia: Loeblioliryloninae Slipinski, 1990 [ ... R] ?/?
 Subfamilia: Ostomopsinae Sen Gupta et Crowson, 1973 [ ... R] ?/?
 Subfamilia: Murmidiinae Jacquelin du Val, 1858 [ ... R] Pal/Rus
 Subfamilia: Ceryloninae Billberg, 1820 [ ... R] Pal/Rus
 =  Aculagnathidae
 =  Lapethinae
 Familia: Alexiidae Imhoff, 1856 [ ... R] Pal/-
 =  Spharosomatidae Ganglbauer, 1899
 Familia: Discolomatidae Horn, 1878 [ ... R] Pal/?
 =  Aphanocephalidae Grouvelle, 1912
 =  Discolomidae
 =  Notiophygidae Jacobson, 1915
 Subfamilia: Notiophiginae Jacobson, 1915 [ ... R] -/-
 Subfamilia: Discolomatinae Horn, 1878 [ ... R] -/-
 Subfamilia: Apanocephalinae Grouvelle, 1912 [ ... R] Pal/?
 Subfamilia: Cephalophaninae John, 1954 [ ... R] -/-
 Subfamilia: Pondonatinae John, 1954 [ ... R] -/-
 Familia: Endomychidae Leach, 1815 [Pg23 ... R] Pal/Rus
 =  Cerasommatidae Bre'thes, 1925
 =  Merophysiidae Seidlitz, 1872
 =  Mycetaeidae Jacquelin du Val, 1858
 Subfamilia: Merophysiinae Seidlitz, 1872 [ ... R] Pal/Rus
 Subfamilia: Holoparamecinae Seidlitz, 1888 [Pg23 ... R] Pal/Rus
 Subfamilia: Leiestinae C.G. Thomson, 1863 [ ... R] Pal/Rus
 Subfamilia: Eupsilobiinae Casey, 1895 [ ... R] Pal/-
 =  Cerasommatidae Bre'thes, 1925
 =  Eidoreinae Sasaji, 1986
 Subfamilia: Endomychinae Leach, 1815 [ ... R] Pal/Rus
 Subfamilia: Epipocinae Gorham, 1873 [ ... R] Pal/Rus
 Subfamilia: Stenotarsinae Chapuis, 1876 [ ... ]
 Subfamilia: Lycoperdininae Redtenbacher, 1844 [Pg23, BalJ ... R] Pal/Rus
 =  Eumorphinae Gistel, 1856
 Subfamilia: Mycetaeinae Jacquelin du Val, 1858 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Anamorphinae Strohecker, 1953 [Pg23, BalJ ... R] ?/?
 =  Mychotheninae Sasaji, 1978
 Subfamilia: Pleganophorinae Jacquelin du Val, 1858 [Pg23 ... R] ?/?
 =  Trochoideinae Chapuis, 1876
 Subfamilia: Xenomycetinae Strohecker, 1962 [ ... R] -/-
 Familia: Coccinellidae Latreille, 1807 [K2cp ... R] Pal/Rus
 =  Epilachnidae
 Subfamilia: Coccidulinae Mulsant, 1846 [ ... ] Pal/Rus
 =  Tetrabrachinae
 Subfamilia: Coccinellinae Latreille, 1807 [Pg31, Flor ... R] Pal/Rus
 Subfamilia: Sticholotidinae Weise, 1901 [Pg23, BalJ ... R] Pal/Rus
 =  Pharinae Casey, 1899
 Subfamilia: Chilocorinae Mulsant, 1846 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Scymninae Mulsant, 1846 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Epilachninae Mulsant, 1846 [ ... R] Pal/Rus
 Familia: Corylophidae LeConte, 1852 [Pg23 ... R] Pal/Rus
 =  Orthoperidae
 Subfamilia: Peltinodinae Paulian, 1950 [ ... R] Pal/-
 Subfamilia: Corylophinae LeConte, 1852 [ ... R] Pal/Rus
 =  Orthoperinae
 Subfamilia: Sericoderinae Matthews, 1888 [ ... R] Pal/Rus
 Subfamilia: Parmulinae Poey, 1854 [ ... R] Pal/Rus
 =  Arthrolipinae
 =  Saciinae
 Familia: Latridiidae Erichson, 1842 [K2sn ... R] Pal/Rus
 =  Lathridiidae
 Subfamilia: Latridiinae Erichson, 1842 [Pg23 ... R] Pal/Rus
 Subfamilia: Corticariinae Curtis, 1829 [K2sn ... R] Pal/Rus
 Superfamilia: Tenebrionoidea Latreille, 1802 [J3ox ... R] Pal/Rus
 =  Heteromera
 Familia: Mycetophagidae Leach, 1815 [Pg23 ... R] Pal/Rus
 =  Tritomidae Crotch, 1873
 Subfamilia: Bergininae Leng, 1920 [ ... R] Pal/?
 Subfamilia: Esarcinae Reitter, 1882 [ ... R] Pal/?
 Subfamilia: Mycetophaginae Leach, 1815 [Pg23 ... R] Pal/Rus
 Familia: Archaeocrypticidae Kaszab, 1964 [ ... R] -/-
 Familia: Pterogeniidae Crowson, 1953 [ ... ]
 Familia: Ciidae Leach, 1819 [Pg23 ... R] Pal/Rus
 =  Cisidae
 =  Octotemnidae
 =  Orophidae
 Subfamilia: Sphindociinae Lawrence, 1974 [ ... R] -/-
 Subfamilia: Ciinae Leach, 1819 [Pg23 ... R] Pal/Rus
 Familia: Tetratomidae Billberg, 1820 [ ... R] Pal/Rus
 Subfamilia: Piseninae Miyatake, 1960 [ ... R] Pal/Rus
 Subfamilia: Tetratominae Billberg, 1820 [ ... R] Pal/Rus
 Subfamilia: Penthinae Lacordaire, 1859 [ ... R] Pal/?
 Familia: Melandryidae Leach, 1815 [K2sn ... R] Pal/Rus
 =  Serropalpidae
 Subfamilia: Hallomeninae Mulsant, 1856/Gistel, 1856 [K2sn, Jant ... R] Pal/Rus
 Subfamilia: Eustrophinae Gistel, 1856 [ ... R] ?/?
 Subfamilia: Melandryinae Leach, 1815 [K2sn, Jant ... R] Pal/Rus
 =  Dircaeinae
 =  Hypulinae
 =  Orchesiinae
 Subfamilia: Osphyinae Mulsant, 1856 [Pg23, BalJ ... R] Pal/Rus
 Familia: Mordellidae Latreille, 1802 [J3ox ... R] Pal/Rus
 =  Liaoximordellidae Wang, 1993
 =  Praemordellidae Schegoleva-Barovskaya, 1929
 Subfamilia: Ctenidiinae Franciscolo, 1951 [ ... R] -/-
 Subfamilia: Mordellinae Latreille, 1802 [J3ox ... R] Pal/Rus
 =  Praemordellinae Schegoleva-Barovskaya, 1929
 Familia: Scraptiidae Mulsant, 1856 [K11 ... R] Pal/Rus
 Familia: Anaspididae Mulsant, 1856 [ ... ] Pal/Rus
 Subfamilia: Scraptiinae Mulsant, 1856 [K11 ... R] Pal/Rus
 Subfamilia: Anaspidinae Mulsant, 1856 [ ... R] Pal/Rus
 Familia: Rhipiphoridae Harold et Gemminger, 1870 [Pg23 ... R] Pal/Rus
 Subfamilia: Pelecotominae Seidlitz, 1875 [Pg23 ... R] Pal/Rus
 Subfamilia: Micholaeminae Vianna, 1971 [ ... R] -/-
 Subfamilia: Ptilophorinae Gerstaecker, 1855 [ ... R] Pal/?
 =  Evaniocerinae
 Subfamilia: Hemirhipidiinae Heller, 1921 [ ... R] -/-
 =  Nephritinae Selander, 1957
 Subfamilia: Rhipidiinae Gerstaecker, 1855 [Pg23 ... R] Pal/Rus
 Subfamilia: Rhipiphorinae Gemminger et Harold, 1870 [Pg23 ... R] Pal/Rus
 Familia: Zopheridae Solier, 1834 [Pg23 ... R] Pal/Rus
 =  Colydiidae Erichson, 1842
 =  Monommatidae Blanchard, 1845
 =  Monommidae
 Subfamilia: Colydiinae Erichson, 1842 [Pg23 ... R] Pal/Rus
 Subfamilia: Pycnomerinae Erichson, 1845 [ ... R] ?/?
 Subfamilia: Zopherinae Solier, 1834 [ ... R] Pal/Rus
 =  Monommatinae Blanchard, 1845
 =  Monomminae
 =  Nosoderminae
 =  Usechinae Horn, 1867
 Familia: Ulodidae Pascoe, 1869 [ ... R] -/-
 =  Mericidae Crowson, 1953
 Familia: Perimylopidae St.George, 1939 [ ... R] -/-
 Familia: Chalcodryidae Watt, 1974 [ ... R] -/-
 Familia: Trachelostenidae Lacordaire, 1859 [ ... ] -/-
 Familia: Tenebrionidae Latreille, 1802 [J3ox ... R] Pal/Rus
 =  Alleculidae Laporte, 1840
 =  Cossiphodidae Wasmann, 1899
 =  Lagriidae Latreille, 1825
 =  Nilionidae Lacordaire, 1859
 =  Petriidae
 =  Rhysopaussidae
 =  Tenthiriidae
 Subfamilia: Lagriinae Latreille, 1825 [Pg23, BalJ ... R] Pal/Rus
 =  Lachninae Billberg, 1820
 =  Pycnocerinae
 Subfamilia: Cossyphinae Latreille, 1802 [ ... ]
 Subfamilia: Nilioninae Lacordaire, 1859 [ ... ]
 Subfamilia: Phrenapatinae Solier, 1834 [ ... R] ?/?
 Subfamilia: Zolodininae Watt, 1974 [ ... R] -/-
 Subfamilia: Pimeliinae Latreille, 1802 [Pg31, Flor ... R] Pal/Rus
 =  Tenthyriinae)
 Subfamilia: Tenebrioninae Latreille, 1802 [Pg23, BalJ ... R] Pal/Rus
 =  Toxicinae
 Subfamilia: Alleculinae Laporte, 1840 [J3ox, KarT ... R] Pal/Rus
 =  Cteniopodidae Solier, 1835
 =  Cistelinae Latreille, 1802
 =  Omophlinae
 =  Xystropodinae Solier, 1835
 Subfamilia: Diaperinae Latreille, 1802 [ ... R] Pal/Rus
 Subfamilia: Coelometopinae Lacordaire, 1859 [ ... R] ?/?
 =  Cnodalinae Gistel, 1856
 =  Stenochiinae Kirby, 1837
 Subfamilia: Cossyphodinae Wasmann, 1899 [ ... R] Pal/-
 Familia: Prostomidae C.G. Thomson, 1859 [ ... R] Pal/Rus
 Familia: Synchroidae Lacordaire, 1859 [Pg31, Flor ... R] ?/?
 Familia: Oedemeridae Latreille, 1810 [Pg23 ... R] Pal/Rus
 Subfamilia: Calopodinae Costa, 1852 [ ... R] Pal/Rus
 Subfamilia: Oedemerinae Latreille, 1810 [Pg23 ... R] Pal/Rus
 Familia: Stenotrachelidae C.G. Thomson, 1859 [ ... R] Pal/Rus
 =  Cephaloidae LeConte, 1862
 =  Nematoplidae LeConte, 1862
 Subfamilia: Stenotrachelinae C.G. Thomson, 1859 [ ... R] Pal/?
 Subfamilia: Nematoplinae LeConte, 1862 [ ... R] Pal/?
 Subfamilia: Cephaloinae LeConte, 1862 [ ... R] Pal/Rus
 Subfamilia: Stolinae Nikitsky, 1985 [ ... R] Pal/-
 Familia: Meloidae Gyllenhal, 1810 [Pg13 ... R] Pal/Rus
 =  Cantharidae Latreille, 1802
 =  Horiidae Latreille, 1802
 =  Tetraonychidae
 Subfamilia: Eleticinae Wellman, 1910 [ ... R] ?/?
 Subfamilia: Meloinae Gyllenhal, 1810 [Pg13 ... R] Pal/Rus
 =  Lyttinae
 =  Mylabrinae
 Subfamilia: Nemognathinae Laporte, 1840 [Pg31 ... R] Pal/Rus
 =  Horiinae
 =  Zonitidinae Mulsant, 1857
 Familia: Mycteridae Blanchard, 1845 [Pg23 ... R] Pal/Rus
 =  Hemipeplidae Lacordaire, 1854
 =  Rhinomaceridae Fleming, 1821
 Subfamilia: Mycterinae Blanchard, 1845 [N13 ... R] Pal/Rus
 Subfamilia: Lacconotinae LeConte, 1862 [ ... R] -/-
 =  Euryphinae J. Thomson, 1860
 Subfamilia: Hemipeplinae Lacordaire, 1854 [Pg23, BalJ ... R] ?/?
 Familia: Boridae C.G. Thomson, 1859 [ ... R] Pal/Rus
 Subfamilia: Borinae C.G. Thomson, 1859 [ ... R] Pal/Rus
 Subfamilia: Synercticinae Lawrence et Pollock, 1994 [ ... R] -/-
 Familia: Trictenotomidae Blanchard, 1845 [ ... R] Pal/-
 Familia: Pythidae Solier, 1834 [K2cm ... R] Pal/Rus
 =  Osphyoplessidae Reitter, 1917
 =?  Ischyomiidae Champion, 1886
 Familia: Pyrochroidae Latreille, 1807 [Pg23 ... R] Pal/Rus
 =  Cononotidae LeConte, 1862
 =  Pedilidae Lacordaire, 1859
 =  Pilipalpidae Abdullah, 1964
 =  Trictenotomidae Crowson, 1980
 Subfamilia: Tydessinae Nikitsky, 1986 [ ... R] Pal/Rus
 Subfamilia: Pilipalpinae Abdullah, 1964 [ ... R] -/-
 Subfamilia: Pedilinae Lacordaire, 1859 [ ... R] Pal/Rus
 Subfamilia: Pyrochroinae Latrielle, 1807 [ ... R] Pal/Rus
 Subfamilia: Agnathinae Lacordaire, 1859 [ ... R] Pal/?
 =  Cononotinae
 Familia: Salpingidae Leach, 1815 [Pg23 ... R] Pal/Rus
 =  Aegialitidae LeConte, 1862
 =  Dacoderidae LeConte, 1862
 =  Elacatidae Chapin, 1923
 =  Eurystethidae Seidlitz, 1916
 =  Inopeplidae Grouvele, 1908
 =  Othniidae LeConte, 1861
 =  Tretothoracidae
 Subfamilia: Othniinae LeConte, 1861 [ ... R] Pal/Rus
 =  Elacatinae
 Subfamilia: Prostominiinae Grouvelle, 1914 [ ... R] ?/?
 =  Trogocryptinae Crowson, 1953
 Subfamilia: Agleninae Horn, 1878 [ ... R] ?/?
 Subfamilia: Salpinginae Leach, 1815 [ ... R] Pal/Rus
 Subfamilia: Inopeplinae Grouvelle, 1908 [ ... R] Pal/?
 Subfamilia: Aegialitinae LeConte, 1862 [ ... R] Pal/Rus
 =  Eurystethinae
 Subfamilia: Dacoderinae LeConte, 1862 [ ... R] -/-
 =  Tretothoracinae
 Familia: Anthicidae Latreille, 1819 [K2-Pg2 ... R] Pal/Rus
 =  Notoxidae Stephens, 1829
 Subfamilia: Eurygeniinae LeConte, 1862 [K2-Pg2 ... R] ?/?
 Subfamilia: Lagriodinae Abdullah et Abdullah, 1968 [ ... R] -/-
 Subfamilia: Afreminae Levey, 1985 [ ... R] -/-
 Subfamilia: Macratriinae LeConte, 1862 [Pg23, BalJ ... R] -/-
 Subfamilia: Steropinae Jacquelin du Val, 1863 [Pg23, BalJ ... R] ?/?
 Subfamilia: Ischaliinae Blair, 1920 [ ... R] ?/?
 Subfamilia: Corobaeninae Abdullah, 1969 [ ... R] -/-
 Subfamilia: Lemodinae Lawrence et Britton, 1991 [ ... R] -/-
 Subfamilia: Tomoderinae Bonadona, 1961 [Pg23, BalJ ... R] ?/?
 Subfamilia: Anthicinae Latreille, 1819 [Pg23, BalJ ... R] Pal/Rus
 =  Notoxinae
 Familia: Aderidae Winkler, 1927 [Pg23 ... R] Pal/Rus
 =  Circeidae Iablokoff-Khnzorian, 1961
 =  Euglenesidae
 =  Euglenidae Seidlitz, 1875
 =  Hylophilidae Pic, 1900
 =  Xylophilidae Shuckard, 1840
 Superfamilia: Chrysomeloidea Latreille, 18?? [ ... ] Pal/Rus
 Familia: Megalopodidae Latreille, 1802 [ ... R] Pal/Rus
 Subfamilia: Palophaginae Kuschel et May, 1990 [ ... R] -/-
 Subfamilia: Zeugophorinae Boeving et Craigheead, 1931 [Pg23 ... R] Pal/Rus
 Subfamilia: Megalopodinae Latreille, 1802 [ ... R] Pal/Rus
 Familia: Orsodacnidae C.G. Thomson, 1859 [ ... R] Pal/Rus
 Subfamilia: Orsodacninae C.G. Thomson, 1859 [ ... R] Pal/Rus
 Subfamilia: Aulacoscelidinae Chapius, 1874 [ ... R] -/-
 Familia: Chrysomelidae Latreille, 1802 [J31 ... R] Pal/Rus
 =  Sagridae Leach, 1815
 Familia: Bruchidae Latreille, 1802 [ ... ] Pal/Rus
 Subfamilia: Bruchinae Latreille, 1802 [Pg31 ... R] Pal/Rus
 =  Pachymerinae Bridwell, 1929
 =  Rhaebinae Chapuis, 1874
 Subfamilia: Hisperinae ??? [ ... R] Pal/Rus
 Subfamilia: Crysomelinae ??? [ ... R] Pal/Rus
 Subfamilia: Eumolpinae ??? [ ... R] Pal/Rus
 Subfamilia: Galerucinae Latreille, 1802 [Pg23 ... R] Pal/Rus
 =  Alticinae
 =  Halticinae Newman, 1834,
 Subfamilia: Donaciinae Kirby, 1837 [Pg1d ... R] Pal/Rus
 Subfamilia: Criocerinae Latreille, 1804 [Pg23 ... R] Pal/Rus
 Subfamilia: Lamprosomatinae Lacordaire, 1848 [ ... ]
 Subfamilia: Cryptocephalinae Gyllenhal, 1813 [Pg13 ... R] Pal/Rus
 =  Chlamisinae
 =  Clytrinae Kirby, 1837
 Familia: Cerambycidae Latreille, 1802 [K12 ... R] Pal/Rus
 =  Hypocephalidae Blanchard, 1845
 =  Parandridae Blanchard, 1845
 =  Spondylidae
 Subfamilia: Parandrinae Blanchard, 1845 [ ... R] Pal/Rus
 Subfamilia: Prioninae Latreille, 1802 [ ... R] Pal/Rus
 Subfamilia: Spondylidinae Audinet-Serville, 1832 [ ... R] Pal/Rus
 Subfamilia: Apatophyseinae Lacordaire, 1869 [ ... R] Pal/?
 Subfamilia: Necydalinae Latreille, 1825 [ ... R] ?/?
 Subfamilia: Lepturinae Latreille, 1802 [ ... R] Pal/Rus
 Subfamilia: Cerambycinae Latreille, 1802 [ ... R] Pal/Rus
 Subfamilia: Lamiinae Latreille, 1825 [ ... R] Pal/Rus
 Familia: Disteniidae J. Thomson, 1860 [ ... R] ?/?
 Familia: Vesperidae Mulsant, 1839 [ ... R] Pal/?
 =  Anoplodermatidae Gue'rin-Me'neville, 1840
 Subfamilia: Anoplodermatinae Gue'rin-Me'neville, 1840 [ ... R] ?/?
 Subfamilia: Philinae J. Thomson, 1860 [N12 ... R] ?/?
 =  Hypocephalinae Blanchard, 1845
 Familia: Oxypeltidae Lacordaire, 1869 [ ... R] -/-
 Infraordo: Curculioniformia Kirejtshuk, 1991 [ ... ] Pal/Rus
 Superfamilia: Curculionoidea [ ... ] Pal/Rus
 =  Rhynchophora Latreille, 1817
+Familia: Obrienidae Zherikhin et Gratshev, 1993 [T2 ... J13] Pal/?
+Subfamilia: Obrieninae Zherikhin et Gratshev, 1993 [??? ... ???] Pal/?
+Subfamilia: Kararhynchinae Zherikhin et Gratshev, 1993 [??? ... ???] Pal/?
+Familia: Ulyanidae Zherikhin, 1993 [K13 ... K13]
 Familia: Nemonychidae Bedel, 1882 [J31 ... R] Pal/Rus
 =  Eubelidae Arnoldi, 1977
 =  Rhinomaceridae Schoenherr, 1832
+Subfamilia: Eobelinae Arnoldi, 1977 [J31 ... ???] Pal/?
+Subfamilia: Brenthorrhininae Arnoldi, 1977 [??? ... ???] Pal/?
+Subfamilia: Nanophydinae Arnoldi, 1977 [??? ... ???] Pal/?
+Subfamilia: Oxycorynoidinae Arnoldi, 1977 [??? ... ???] Pal/?
 Subfamilia: Nemonychinae Bedel, 1882 [ ... R] Pal/Rus
 Subfamilia: Rhinorhynchinae Voss, 1922 [ ... R] -/-
 Subfamilia: Doydirhynchinae Pierce, 1916 [ ... R] Pal/-
 =  Cimberidinae Gozis, 1882
 =  Neocimberidinae
+Subfamilia: Slonikinae Zherikhin, 1977 [ ... ]
 Familia: Anthribidae Billberg, 1820 [K1 ... R] Pal/Rus
 =  Bruchelidae
 =  Platyrhynidae
 =  Platystomatidae
 =  Platystomidae
 =  Urodontidae
 Subfamilia: Urodontinae C.G. Thomson, 1859 [ ... R] Pal/Rus
 =  Bruchelinae
 Subfamilia: Anthribinae Billberg, 1820 [ ... R] Pal/Rus
 =  Brachytarsinae
 Subfamilia: Choraginae Kirby, 1819 [ ... R] ?/?
 =  Araeocerinae
+Familia: Eccortarthridae Arnoldi, 1977 [J?? ... ???] Pal/?
+Subfamilia: Eccortarthrinae Arnoldi, 1977 [J?? ... ???] Pal/?
+Subfamilia: Baissorhynchinae Zherikhin, 1993 [ ... ] Pal/?
 Familia: Belidae Schoenherr, 1826 [ ... R] ?/?
 Subfamilia: Belinae Schoenherr, 1826 [ ... R] ?/?
 Subfamilia: Pachyurinae Kuschel, 1959 [ ... R] -/-
 Familia: Oxycorynidae Schoenherr, 1840 [ ... R] ?/?
 Subfamilia: Oxycoryninae Schoenherr, 1840 [ ... R] ?/?
 Subfamilia: Afrocoryninae Voss, 1957 [ ... R] -/-
 Subfamilia: Aglycyderinae Wollaston, 1864 [ ... R] -/-
 =  Proterhininae
 Subfamilia: Allocoryninae Sharp, 1890 [ ... R] ?/?
 Subfamilia: Hispodinae Voss, 1957 [ ... R] ?/?
 Subfamilia: Metrioxeninae Voss, 1953 [ ... R] ?/?
 Familia: Rhynchitidae Gistel, 1865 [ ... R] Pal/Rus
 =  Pterocolidae Lacordaire, 1866
 Subfamilia: Rhynchitinae Gistel, 1865 [ ... R] Pal/Rus
 Subfamilia: Pterocolinae Lacordaire, 1866 [ ... R] ?/?
+Subfamilia: Isotheinae Scudder, 1893 [??? ... ???] ?/?
 Familia: Attelabidae Billberg, 1820 [K12 ... R] Pal/Rus
 =  Apoderidae Jekel, 1860
 Subfamilia: Attelabinae Billberg, 1820 [ ... R] Pal/Rus
 Subfamilia: Apoderinae Jekel, 1860 [ ... R] Pal/Rus
 Subfamilia: Archolabinae Voss, 1929 [ ... R] ?/?
 Familia: Brentidae Billberg, 1820 [Pg33 ... R] Pal/Rus
 =  Brenthidae
 =  Cyladidae
 =  Eurhynchidae
 Subfamilia: Brentinae Billberg, 1820 [ ... R] Pal/Rus
 Subfamilia: Antliarhininae Schoenherr, 1823 [ ... R] ?/?
 Subfamilia: Cyladinae [ ... R] ?/?
 Subfamilia: Cyphagoginae Kolbe, 1892 [ ... R] ?/?
 Subfamilia: Pholidochlamidinae Damoiseau, 1962 [ ... R] ?/?
 Subfamilia: Taphroderinae Lacordaire, 1866 [ ... R] ?/?
 Subfamilia: Trachelizinae Lacordaire, 1866 [ ... R] ?/?
 Subfamilia: Ulocerinae Schoenherr, 1823 [ ... R] ?/?
 Subfamilia: Eurhynchinae Lacordaire, 1863 [ ... R] ?/?
 Familia: Apionidae Schoenherr, 1823 [ ... R] Pal/Rus
 Subfamilia: Apioninae Schoenherr, 1823 [ ... R] Pal/Rus
 Subfamilia: Tanainae [ ... R] ?/?
 Subfamilia: Myrmacicelinae Zimmerman, 1994 [ ... R] -/-
 Subfamilia: Rhinorhynchidiinae Zimmerman, 1994 [ ... R] -/-
 Subfamilia: Nanophyinae Gistel, 1856 [ ... R] Pal/Rus
 Familia: Caridae Thomson, 1992 [ ... R] -/-
 Familia: Ithyceridae Schoenherr, 1823 [ ... R] -/-
 Familia: Curculionidae Latreille, 1802 [K13 ... R] Pal/Rus
 =  Calendridae
 =  Cossonidae Schoenherr, 1825
 =  Platypodidae
 =  Rhynchophoridae Schoenherr, 1833
 =  Scolytidae
 Subfamilia: Brachycerinae Billberg, 1820 [ ... R] Pal/Rus
 =  Alophinae LeConte, 1874
 =  Amycterinae G.R.Waterhouse, 1854
 =  Aterpinae Lacordaire, 1863
 =  Brachyderinae Schoenherr, 1826
 =  Byrsopinae Germar, 1829
 =  Cryptolaringinae Schalkwyk, 1966
 =  Cylindrorhininae
 =  Desmidophorinae Morimoto, 1962
 =  Diabathrariinae Lacordaire, 1863
 =  Dinomorphinae Lacordaire, 1863
 =  Ectemnorhininae Lacordaire, 1863
 =  Entiminae Schoenherr, 1823
 =  Eremninae
 =  Goniopterinae Lacordaire, 1863
 =  Hipporhininae
 =  Hyperinae Lacordaire, 1863
 =  Leptopiinae
 =  Microcerinae Lacordaire, 1863
 =  Ocladiinae Lacordaire, 1866
 =  Otiorhynchinae Schoenherr, 1826
 =  Pachyrhynchinae Schoenherr, 1826
 =  Rhadinosominae
 =  Rhyparosominae
 =  Rhytirhininae Lacordaire, 1863
 =  Somatodinae Lacordaire, 1863
 =  Tanyrhynchinae Schoenherr, 1826
 =  Thecesterninae Lacordaire, 1863
 =  Ulomascinae Lacordaire, 1866
 =  Viticinae Morimoto, 1983
 Subfamilia: Curculioninae Latreille, 1802 [ ... R] Pal/Rus
 =  Anoplinae Bedel, 1884
 =  Anthonominae C.G.Thomson, 1859
 =  Bagoinae C.G.Thomson, 1859
 =  Baridinae Schoenherr, 1836
 =  Camarotinae Schoenherr, 1833
 =  Campyloscelinae Schoenherr, 1845
 =  Ceratopodinae Lacordaire, 1863
 =  Ceutorhynchinae Gistel, 1856
 =  Cioninae Schoenherr, 1825
 =  Cleoninae Schoenherr, 1826
 =  Cryptorhynchinae Schoenherr, 1825 ?/?
 =  Erirhininae Schoenherr, 1825
 =  Eugnominae Lacordaire, 1863
 =  Tadiinae Zimmerman, 1993
 Subfamilia: Gymnetrinae [ ... ]
 =  Lixinae Schoenherr, 1823
 =  Magdalininae Pascoe, 1870
 =  Molytinae Schoenherr, 1823
 =  Nerthopinae Lacordaire, 1866
 =  Otidocephalinae Lacordaire, 1863
 Subfamilia: Prionomerinae [ ... ]
 =  Raymondionymidae Reitter, 1913
 =  Myrtonyminae Kuschel, 1990
 =  Pyropinae Lacordaire, 1866
 =  Rhyncheninae
 =  Tachigoninae
 =  Trigonocolinae Lacordaire, 1863
 =  Tychiinae C.G.Thomson, 1859
 =  Zygopinae Lacordaire, 1866
 Subfamilia: Brachyceropseinae Aurivillius, 1926 [ ... R] ?/?
 Subfamilia: Conoderinae Schoenherr, 1833 [ ... R] ?/?
 Subfamilia: Cyclominae Schoenherr, 1826 [ ... R] ?/?
 Subfamilia: Mesoptiliinae Lacordaire, 1863 [ ... R] ?/?
 Subfamilia: Orobitidinae C.G.Thomson, 1859 [ ... R] ?/?
 Subfamilia: Xiphaspidinae Marshall, 1920 [ ... R] ?/?
 Subfamilia: Dryophthorinae Schoenherr, 1825 [ ... R] ?/?
 =  Calandrinae
 =  Calendrinae
 =  Cryptodermatinae Bovie, 1908
 =  Orthognathinae Lacordaire, 1866
 =  Rhynchophorinae Schoenherr, 1833
 =  Stromboscerinae Lacordaire, 1866
 Subfamilia: Cossoninae Schoenherr, 1825 [ ... R] Pal/Rus
 Subfamilia: Scolytinae Latreille, 1804 [ ... R] Pal/Rus
 =  Coptonotinae
 =  Hylesininae
 =  Ipinae
 =  Tomicinae
 Subfamilia: Platypodinae Shuckard, 1840 [ ... R] Pal/Rus
 =  Chapuisiinae
 =  Crossotarsinae
 =  Diaporinae
 =  Periommatinae
 =  Platytarsilinae
 Subordo: Myxophaga Crowson, 1955 [ ... ] Pal/Rus
 Infraordo: Microsporomorpha ???Crotch, 1873 [ ... ] Pal/Rus
+Superfamilia: Asiocoleoidea Rohdendorf, 1961 [ ... ]
+Familia: Asiocoleidae Rohdendorf, 1961 [P2uf ... P2kz] Pal/?
+Familia: Tricoleidae Ponomarenko, 1969 [T2l ?T3r ... T3] Pal/?
+Superfamilia: Rhombocoleoidea Rohdendorf, 1961 [ ... ]
+Familia: Rhombocoleidae Rohdendorf, 1961 [P2uf ... P2t] Pal/?
+Superfamilia: Schizophoroidea Ponomarenko, 1968 [ ... ]
+Familia: Schizophoridae Ponomarenko, 1968 [P2kz ... K1 (?K2)] Pal/?
+Familia: Catiniidae Ponomarenko, 1968 [T2l ... J31] Pal/?
+Familia: Schizocoleidae Rohdendorf, 1961 [P2kz, Suri ... K1ap, Bon] Pal/?
 =  Curculiopseidae Martynov, 1937
 Superfamilia: Microsporoidea Crotch, 1873 [ ... R]
 Familia: Lepiceridae Hinton, 1936 [ ... R] -/-
 =  Cyathoceridae Scharp, 1882
 Familia: Microsporidae Crotch, 1873 [ ... R] Pal/Rus
 =  Sphaeriidae Erichson, 1845
 Familia: Hydroscaphidae LeConte, 1874 [ ... R] Pal/?
 Familia: Torridincolidae Steffan, 1964 [ ... R] Pal/?
 Subordo: Adephaga Schellenberg, 1806 [ ... ] Pal/Rus
 Infraordo: Haliplomorpha Brulle', 1835 [ ... ] Pal/Rus
 Superfamilia: Triaploidea Ponomarenko, 1977 [ ... ]
 Familia: Triaplidae Ponomarenko, 1977 [T2l ... T3c] Pal/?
 Superfamilia: Haliploidea [ ... ] Pal/Rus
 Familia: Haliplidae Brulle', 1835 [Kal3-cm1 ... R] Pal/Rus
 Infraordo: Carabomorpha [ ... ] Pal/Rus
 Superfamilia: Gyrinoidea ???Latreille, 1810 [ ... ] Pal/Rus
 Familia: Gyrinidae Latreille, 1810 [J11 ... R] Pal/Rus
 Subfamilia: Spanglerogyrininae Folkerts, 1979 [ ... R] -/-
 Subfamilia: Gyrininae Latreille, 1810 [J11 ... R] Pal/Rus
 =  Orectochilinae
 Superfamilia: Dytiscoidea Leach, 1815 [ ... ] Pal/Rus
 Familia: Colymbothetidae Ponomarenko, 1993 [T3r ... ???] Pal/?
 Familia: Hygrobiidae Regimbart, 1878 [N1aq ... R] Pal/?
 =  Pelobiidae
 Familia: Parahygrobiidae Ponomarenko, 1977 [J31 ... ???] Pal/?
 Familia: Coptoclavidae Ponomarenko, 1961 [J11 ... K1ap] Pal/?
 Subfamilia: Timarchopsinae [J11 ... K1ha] Pal/?
 =  Necronectinae Ponomarenko, 1977
 Subfamilia: Charonoscaphinae Ponomarenko, 1977 [J3ox, KarT ... ???] Pal/?
 Familia: Liadytidae Ponomarenko, 1977 [J11 ... J3] Pal/?
 Familia: Noteridae Thomson, 1860 [Pg12 ... R] Pal/Rus
 =  Phreatodytidae Ueno, 1957
 Familia: Noteridae Thomson, 1860 [Pg12 ... R] Pal/Rus
 Familia: Phreatodytidae Ueno, 1957 [ ... R] Pal/-
 Familia: Amphizoidae Lacordaire, 1854 [ ... R] Pal/?
 Familia: Dytiscidae Leach, 1815 [J31 ... R] Pal/Rus
 Subfamilia: Palaeogyrininae Hatch, 1927 [Pg12 ... N1aq, Rot] ?/?
 Subfamilia: Copelatinae Van den Branden, 1885 [N12, Stav ... R] Pal/Rus
 Subfamilia: Aubehydrinae Guignot, 1942 [ ... R] ?/?
 Subfamilia: Laccophilinae Gistel, 1856 [Pg22, GreR ... R] Pal/Rus
 Subfamilia: Hydroporinae Sharp, 1880 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Colymbetinae Erichson, 1837 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Dytiscinae Leach, 1815 [Pg31, Flor ... R] Pal/Rus
 Superfamilia: Caraboidea ???Latreille, 1802 [T2l ... R] Pal/Rus
 Familia: Trachypachidae Thomson, 1857 [T2l ... R] Pal/Rus
 =  Leptopodocoleidae Hong, 1982
 Subfamilia: Trachypachinae Thomson, 1857 [T2l ... R] Pal/Rus
 Subfamilia: Eodromeinae Ponomarenko, 1977 [T2-3l-ca ... K1ap] Pal/?
 =  Leptopodocoleinae Hong, 1982
 Familia: Carabidae Latreille, 18O2 [J11 ... R] Pal/Rus
 =  Cicindelidae
 =  Paussidae
 Subfamilia: Protorabinae Ponomarenko, 1977 [J1he ... K1al] Pal/?
 Subfamilia: Gehringiinae Darlington, 1933 [ ... R] ?/?
 Subfamilia: Cicindelinae Latreille, 1802 [Pg22 ... R] Pal/Rus
 Subfamilia: Paussinae Latreille, 1807 [Pg23 ... R] Pal/Rus
 Subfamilia: Nototylinae Reichardt, 1977 [ ... R] ?/?
 Subfamilia: Cicindinae Baenninger, 1927 [ ... R] Pal/Rus
 Subfamilia: Metriinae Leconte, 18?? [Pg23, BalJR ... R]
 Subfamilia: Omophroninae Bonelli, 1810 [N13 ... R] Pal/Rus
 Subfamilia: Carabinae Latreille, 1802 [K2tu ... R] Pal/Rus
 Subfamilia: Hiletinae Schioedte, 1847 [ ... R] ?/?
 Subfamilia: Loricerinae Bonelli, 1810 [ ... R] ?/?
 Subfamilia: Elaphrinae Latreille, 1802 [ ... R] Pal/Rus
 Subfamilia: Migadopinae Chadoir, 1861 [ ... R] ?/?
 Subfamilia: Siagoninae Bonelli, 1810 [ ... R] ?/?
 Subfamilia: Scaritinae Bonelli, 1810 [ ... R] Pal/Rus
 Subfamilia: Trechinae Bonelli, 1810 [ ... R] Pal/Rus
 Subfamilia: Harpalinae Bonelli, 1810 [ ... R] Pal/Rus
 Subfamilia: Pseudomorphinae Newmann, 1842 [ ... R] -/-
 Subfamilia: Brachininae Bonelli, 1810 [ ... R] Pal/Rus
 Superfamilia: Rhysodoidea ???Laporte, 1840 [Pg33 ... R] Pal/Rus
 Familia: Rhysodidae Laporte, 1840 [Pg33 ... R] Pal/Rus