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Tolotra Ranarilalatiana, Lala Harivelo Raveloson Ravaomanarivo, Johannes Bergsten. 2019.
Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non-erichsonii group species.
ZooKeys, 2019, 869: 19-90.
doi: 10.3897/zookeys.869.33997
http://zoobank.org/B7C88A64-C06E-4B67-A352-F2F9C8FB0D1C