Жужелицы находятся во многих отношениях в более выгодном положении, чем другие крупные семейства жуков (по крайней мере -- в России). Во-первых, они являются излюбленным объектом и профессионалов, и любителей -- их изучает очень много специалистов. Во-вторых, жужелицы территории бывшего СССР обеспечены великолепной базой данных, которую создал и много лет профессионально поддерживает Кирилл Макаров. О состоянии и возможностях этой базы данных красноречиво говорит тот факт, что из нее полностью программным способом (без всяких последующих поправок и редактирования) был получен оригинал-макет самого лучшего каталога жуков, из когда либо изданных в России -- Kryzhanovskij O.L., Belousov I.A., Kabak I.I., Kataev B.M., Makarov K.V., Shilenkov V.G., 1995. A Checklist of the ground-beetles of Russia and adjacent Lands (Insecta, Coleoptera, Carabidae) [Pensoft, Series Faunistica, 3: 271 рр.]
К.В. Макаров любезно предоставил эту уникальную базу данных для использования на нашем сайте. Пока ряд технических причин не позволяет нам организовать интерактивный поиск в базе данных. Поэтому мы можем предложить вашему вниманию только отдельные выборки, которые не превышают по объему 100-200 килобайт. Первым мы помещаем общий обзор системы семейства Carabidae, принятой в базе данных. В выборку включены все таксоны с рангами от семейства до подрода и их синонимы.
А. Лобанов, март 2000 г.
Список сгенерирован из базы данных CARABIDA
(К.В. Макаров, версия от 21 марта 2000 г.)
Familia: CARABIDAE Subfamilia: CICINDELINAE Supertribus: CICINDELITAE Tribus: MEGACEPHALINI Subtribus: MEGACEPHALINA Genus: Megacephala Latr., 1802 = Tetracha Hope, 1838 Subgenus: Grammognatha Motsch., 1850 Tribus: CICINDELINI Subtribus: CICINDELINA Genus: Cicindela L., 1758 Subgenus: Eumecus Motsch., 1850 = Cylindera Westw., 1831 = Cylindrodera Bed., 1879 = Cylindella Jacobs., 1924 Subgenus: Myriochile Motsch., 1862 = Monelica Rivalier, 1950 = Catopria Guerin, 1849 = Lutaria Horn, 1891 Subgenus: Cephalota Dokht., 1883 = Taenidia Rivalier, 1950 = Spiralia Rivalier, 1950 = Cassolaia Wiesner, 1985 Subgenus: Cicindina Adam & Merkl, 1986 = Eugrapha Rivalier, 1950 Subgenus: Lophyridia Jeann., 1946 Subgenus: Laphyra Dej., 1836 = Lophyra Motsch., 1859 = Tribonophora Rivalier, 1950 = Chaetostyla Ganglb., 1892 = Chaetodera Jeann., 1946 Subgenus: Cicindela L., 1758 Subfamilia: OMOPHRONINAE Tribus: OMOPHRONINI Genus: Omophron Latr., 1802 = Epactius Schneider, 1791 = Scolytus F., 1791 Subgenus: Omophron Latr., 1802 Subfamilia: CARABINAE Supertribus: NEBRIITAE Tribus: NEBRIINI Genus: Pelophila Dej., 1826 Genus: Leistus Froel., 1799 Subgenus: Pogonophorus Latr., 1802 = Oreobius Dan., 1903 = Eurynophorus Breit, 1914 = Chaetoleistus Sem., 1904 Subgenus: Oreoleistus Kryzh. & Shil., in litt Subgenus: Leistus Froel., 1799 = Leistidius Dan., 1903 = Leistophorus Rtt., 1905 = Acroleistus Rtt., 1905 = Euleistulus Rtt., 1905 = Neoleistus Erwin, 1970 Genus: Nebria Latr., 1825 Subgenus: Eunebria Jeann., 1937 Subgenus: Paranebria Jeann., 1937 Subgenus: Boreonebria Jeann., 1937 Subgenus: Eonebria Sem., 1928 Subgenus: Orientonebria Shil., 1975 Subgenus: Reductonebria Shil., 1975 Subgenus: Catonebria Shil., 1975 Subgenus: Nebria Latr., 1825 = Helobia Steph., 1828 Subgenus: Alpaeus Bon., 1809 = Alpaeonebria Csiki, 1946 Supertribus: NOTIOPHILITAE Tribus: NOTIOPHILINI Genus: Notiophilus Dum., 1806 Subgenus: Notiophilus Dum., 1806 Subgenus: Latviaphilus Barsevskis, 1994 Subgenus: Makarovius Barsevskis, 1994 Supertribus: CARABITAE Tribus: CARABINI Genus: Calosoma Web., 1801 Subgenus: Calosoma Web., 1801 = Callipara Motsch., 1865 = Calipera Bed., 1881 = Syncalosoma Breun., 1927 Subgenus: Acalosoma Lafer, 1990 Subgenus: Campalita Motsch., 1865 = Cosmoplata Motsch., 1865 = Callistrata Motsch., 1865 = Catasoma Lap., 1929 = Eremosoma Lap., 1929 Subgenus: Caminara Motsch., 1865 Subgenus: Charmosta Motsch., 1865 Genus: Callisthenes Fisch., 1821 Subgenus: Callisthenes Fisch., 1821 Subgenus: Callisphaena Motsch., 1865 Genus: Carabus L., 1758 = Tachypus Web., 1801 Subgenus: Acrocarabus Lap., 1931 Subgenus: Eucarabus Geh., 1885 = Eutelocarabus Geh., 1885 = Xystrocarabus Rtt., 1896 = Tylocarabus Rtt., 1896 = Araeocarabus Rtt., 1896 = Apocarabus Lap., 1930 = Parhomopterus Lap., 1931 = Orientocarabus Kwon & Lee, 1983 Subgenus: Autocarabus Seidl., 1887 = Goniocarabus Rtt., 1896 = Loxocarabus Rtt., 1896 = Cancellocarabus Lutsh., 1924 Subgenus: Carabus L., 1758 = Carabus Thoms., 1875 = Paracarabus Lap., 1931 = Neocarabus Hatch, 1949 Subgenus: Morphocarabus Geh., 1885 = Apostocarabus Rtt., 1896 = Pancarabus Rtt., 1896 = Promorphocarabus Rtt., 1896 = Ancylocarabus Rtt., 1896 = Gigantocarabus Sem., 1898 = Sajanocarabus Lutsh., 1924 = Amorphocarabus Lap., 1930 = Basilicarabus Lap., 1930 Subgenus: Leptinocarabus Rtt., 1896 Subgenus: Trachycarabus Geh., 1885 = Lyperocarabus Lap., 1930 Subgenus: Ophiocarabus Rtt., 1896 = Haplocarabus Sem., 1898 Subgenus: Cryptocarabus Rtt., 1896 = Microcarabus Ishikawa, 1979 Subgenus: Cyclocarabus Rtt., 1896 Subgenus: Mimocarabus Geh., 1885 Subgenus: Archicarabus Seidl., 1887 = Deuterocarabus Rtt., 1896 = Aptocarabus Rtt., 1896 = Rhipocarabus Rtt., 1896 = Nemoralus Schuler, 1976 Subgenus: Limnocarabus Geh., 1885 Subgenus: Homoeocarabus Rtt., 1896 Subgenus: Hemicarabus Geh., 1885 Subgenus: Aulonocarabus Rtt., 1896 Subgenus: Leptocarabus Geh., 1885 = Adelocarabus Rtt., 1896 Subgenus: Asthenocarabus Lap., 1931 Subgenus: Diocarabus Rtt., 1896 = Allocarabus Lap., 1930 Subgenus: Pachycarabus Geh., 1876 = Caucasocarabus Lap., 1931 Subgenus: Orinocarabus Kr., 1878 = Carpatophilus Rtt., 1896 Subgenus: Hadrocarabus Thoms., 1875 = Mesocarabus Thoms., 1875 Subgenus: Oreocarabus Geh., 1876 = Euporocarabus Rtt., 1896 = Phricocarabus Rtt., 1896 = Piocarabus Rtt., 1896 = Cytilocarabus Rtt., 1896 Subgenus: Ulocarabus Rtt., 1896 = Bactrocarabus Sem., 1927 Subgenus: Semnocarabus Rtt., 1896 = Anthracocarabus Lap., 1930 = Zoocarabus Rtt., 1896 Subgenus: Tomocarabus Rtt., 1896 = Callistocarabus Rtt., 1896 = Procrustides Sem., 1905 = Eurycarabus sensu Breun., 1932 Subgenus: Scambocarabus Rtt., 1896 = Eremocarabus Lap., 1931 Subgenus: Pachystus Motsch., 1865 = Melancarabus Thoms., 1875 Subgenus: Hygrocarabus Thoms., 1875 Subgenus: Chaetocarabus Thoms., 1875 Subgenus: Platycarabus Mor., 1886 = Pseudocechenus Mor., 1886 = Baudiicarabus Ganglb., 1909 Subgenus: Panthophyrtus Thieme, 1881 Subgenus: Megodontus Sol., 1848 = Proteocarabus Geh., 1876 = Aulacocarabus Geh., 1876 = Nabicarabus Kwon & Lee, 1983 Subgenus: Ainocarabus Mandl, 1973 Subgenus: Pachycranion Sol., 1848 = Pachycranius Thoms., 1875 Subgenus: Carabulus Lutsh., 1924 Subgenus: Chrysocarabus Thoms., 1875 Subgenus: Acoptolabrus Mor., 1886 = Koreacoptolabrus Kwon & Lee, 1989 = Neodamaster Nakajima, 1993 Subgenus: Sphodristocarabus Geh., 1885 = Sphodristus Thoms., 1875 Subgenus: Cechenochilus Motsch., 1850 = Cechenus Fisch., 1822 Subgenus: Eotribax Sem., 1898 Subgenus: Leptoplesius Rtt., 1898 Subgenus: Cechenotribax Sem. & Zn., 1932 Subgenus: Cratocechenus Rtt., 1896 Subgenus: Cratocephalus Kirsch, 1859 = Pachycechenus Sem., 1898 Subgenus: Pseudotribax Kr., 1884 Subgenus: Cratophyrtus Rtt., 1896 Subgenus: Cratocarabus Rtt., 1896 Subgenus: Alipaster Rtt., 1896 Subgenus: Tribax Fisch., 1817 = Platychrus Kol., 1845 Subgenus: Microplectes Rtt., 1896 Subgenus: Microtribax Gottw., 1982 = Plectes Fisch., 1832 = Neoplectes Rtt., 1885 Subgenus: Archiplectes Gottw., 1982 Subgenus: Neoplectes Gottw., 1982 = Plectocarabus Deuve, 1991 Subgenus: Lamprostus Motsch., 1865 = Chaetogaster Lap., 1931 = Chaetoprostus Rtt., 1896 = Pseudoprocrustes Mor., 1886 Subgenus: Procrustes Bon., 1809 = Macrogenus Motsch., 1846 Subgenus: Goniocarabus Geh., 1885 = Alagocarabus Mor., 1886 = Acarabus Sem., 1890 = Goniognathus Kr., 1883 Subgenus: Deroplectes Rtt., 1895 Subgenus: Plesius Sem., 1905 = Paraplesius Mor., 1886 Subgenus: Axinocarabus Mor., 1886 Subgenus: Coptolabrus Sol., 1848 = Eocarabus Sem., 1898 = Coptolabrulus Rtt., 1897 = Eucoptolabrus Sem., 1898 = Nescocoptolabrus Lap., 1930 Subgenus: Damaster Koll., 1836 = Adamaster Rtt., 1896 Subgenus: Procerus Dej., 1826 Subgenus: Eupachys Chaud., 1857 Subgenus: Lipaster Motsch., 1865 = Lamprocarabus Thoms., 1875 = Titanocechenus Breun. & Ruspoli, 1970 Tribus: CYCHRINI Genus: Cychrus F., 1774 Supertribus: ELAPHRITAE Tribus: ELAPHRINI Genus: Diacheila Motsch., 1845 = Diachila auct. = Arctobia Thoms., 1859 Genus: Blethisa Bon., 1810 = Helobium Leach, 1815 = Rhaphiona Fisch., 1825 Genus: Elaphrus F., 1775 Subgenus: Arctelaphrus Sem., 1926 Subgenus: Neoelaphrus Hatch, 1951 Subgenus: Elaphrus F., 1775 Subgenus: Elaphroterus Sem., 1895 = Elaphrotatus Sem., 1895 Supertribus: LORICERITAE Tribus: LORICERINI Genus: Loricera Latr., 1802 = Lorocera auct. Subgenus: Loricera Latr., 1802 Supertribus: SIAGONITAE Tribus: SIAGONINI Genus: Siagona Latr., 1804 Tribus: CYMBIONOTINI Genus: Cymbionotum Baudi, 1864 = Graniger auct., nec Motsch., 1864 = Coscinia Dej., 1831 Supertribus: SCARITITAE Tribus: SCARITINI Genus: Scarites F., 1775 Subgenus: Distichus Motsch., 1857 = Lophogenius Motsch., 1857 = Scaritodes Chaud., 1879 = Adialampus Goz., 1882 = Dischistus Portevin, 1929 Subgenus: Scarites F., 1775 = Parallelomorphus Motsch., 1849 = Harpalites Motsch., 1850 = Broscomorphus Motsch., 1857 = Paramecomorphus Motsch., 1857 Subgenus: Scallophorites Motsch., 1857 Tribus: CLIVININI Genus: Clivina Latr., 1802 = Ceratoglossa MacLeay, 1866 Genus: Coryza Putz., 1866 Genus: Orientoreicheia Bulirsch & Hurka, 1994 Tribus: DYSCHIRIINI Genus: Dyschirius Bon., 1810 = Dyschiridius Jeann., 1941 Genus: Dyschiriodes Jeann., 1941 = Dyschirius Jeann., 1941 Subgenus: Eudyschirius Fed., in litt Subgenus: Dyschiriodes Fed., in litt Subgenus: Chiridysus Fed., in litt Subgenus: Paradyschirius Fed., in litt Genus: Reicheiodes Ganglb., 1892 Subgenus: Reicheiodes Ganglb., 1892 Genus: Clivinopsis Bed., 1895 = Clivinaxis Rtt., 1909 Supertribus: BROSCITAE Tribus: BROSCINI Genus: Broscus Panz., 1813 = Cephalotes Bon., 1811 = Pseudocupius Voet Genus: Eobroscus Kryzh., 1951 Genus: Craspedonotus Schaum, 1863 = Pseudobroscus Sem., 1888 Genus: Miscodera Esch., 1830 = Leiochiton Curt., 1831 Genus: Broscosoma Putz., 1846 Tribus: APOTOMINI Genus: Apotomus Ill., 1807 Supertribus: TRECHITAE Tribus: TRECHINI Subtribus: PERILEPTINA Genus: Perileptus Schaum, 1860 = Ochthephilus Nietner, 1857 Genus: Neoblemus Jeann., 1923 Subtribus: TRECHODINA Genus: Thalassophilus Woll., 1854 Genus: Eotrechodes Ueno, Lafer & Sunduk., 1995 Subtribus: TRECHINA Genus: Aepiblemus Bel. & Kabak, 1993 Genus: Trechoblemus Ganglb., 1896 Genus: Oroblemites Ueno & Pawl., 1981 Genus: Pseudanophthalmus Jeann., 1920 = Duvaliopsis Jeann., 1928 = Tenessarius Val., 1952 = Aphanotrechus Barr, 1960 Genus: Blemus Dej., 1821 = Lasiotrechus Ganglb., 1892 Genus: Nannotrechus Winkl., 1926 = Birsteiniotrechus Ljov., 1972 Subgenus: Nannotrechus Winkl., 1926 Subgenus: Archinannotrechus Bel., 1998 Subgenus: Abasorites Bel., 1998 Genus: Alanorites Bel., in litt Genus: Cimmerites Jeann., 1928 Genus: Paracimmerites Bel., 1998 Genus: Taurocimmerites Bel., 1998 Genus: Porocimmerites Bel., 1998 Genus: Troglocimmerites Ljov., 1970 Genus: Apocimmerites Bel., 1998 Genus: Taniatrechus Bel. & Dolzh., 1994 Genus: Jeannelius Kurn., 1959 Genus: Pseudaphaenops Winkl., 1914 Genus: Caucasaphaenops Bel., in litt Genus: Meganophtalmus Kurn., 1959 Genus: Caucasorites Bel. & Zamot., 1997 Genus: Duvalius Delar., 1859 Subgenus: Duvalius Delar., 1859 Subgenus: Duvalidius Jeann., 1928 Subgenus: Duvaliotes Jeann., 1928 Genus: Inotrechus Dolzh. & Ljov., 1989 Genus: Masuzoa Ueno, 1960 Genus: Epaphiopsis Ueno, 1953 = Glabrotrechus Suenson, 1957 Subgenus: Epaphiama Jeann., 1962 Genus: Trechiama Jeann., 1927 Subgenus: Leptepaphiama Jeann., 1962 Genus: Epaphius Steph., 1827 Genus: Trechus Clairv., 1806 = Blemus Steph., 1830 = Calotrechus Woll., 1854 = Altaiotrechus Khnz., 1971 Tribus: TACHYINI = LIMNASTINI Genus: Tachys Steph., 1829 Subgenus: Tachys Steph., 1829 = Isotachys Casey, 1918 = Macrotachys Kult, 1961 Subgenus: Paratachys Casey, 1918 = Eotachys Jeann., 1941 Genus: Polyderis Motsch., 1862 = Microtachys Casey, 1918 = Neotachys Kult, 1961 = Polyderidius Jeann., 1962 Genus: Elaphropus Motsch., 1839 = Tachylopha Motsch., 1862 = Barytachys Chaud., 1868 = Trepanotachys Alluaud, 1933 = Tachyphanes Jeann., 1946 Subgenus: Elaphropus Motsch., 1839 Subgenus: Tachyura Motsch., 1862 Subgenus: Amaurotachys Jeann., 1946 Subgenus: Sphaerotachys J.Mull., 1926 Genus: Porotachys Net., 1914 Genus: Tachyta Kirby, 1837 = Tachymenis Motsch., 1857 Genus: Lymnastis Motsch., 1862 = Limnastis auct. = Zuphiolum Fairm., 1896 = Paralimnastis Jeann., 1936 Tribus: BEMBIDIINI Genus: Asaphidion Goz., 1886 = Tachypus Dej., 1821 Genus: Ocys Steph., 1829 Subgenus: Ocys Steph., 1829 Subgenus: Oreocys Peyerimhoff, 1908 Genus: Cillenus Sam., 1819 = Cillenum Dej., 1821 Subgenus: Sakagutia Ueno, 1955 Genus: Bembidion Latr., 1802 = Bembidium auct. = Bembicidium Gemm. & Har., 1868 Subgenus: Desarmatocillenus Net., 1942 Subgenus: Microserrullula Net., 1921 = Serrula Net., 1910 Subgenus: Bracteon Bed., 1879 = Chrysobracteon Net., 1914 = Parabracteon Notman, 1929 = Argyrobracteon Net., 1942 = Conicibracteon Net., 1942 = Foveobracteon Net., 1942 = Litoreobracteon Net., 1942 = Stylobracteon Net., 1942 Subgenus: Odontium Lec., 1848 = Cylindrobracten Net., 1939 Subgenus: Eurytrachelus Motsch., 1850 = Eudromus Kirby, 1837 = Trachypachus Motsch., 1844 = Platytrachelus Motsch., 1844 = Pogonidium Ganglb., 1892 = Hydrium Jeann., 1941 Subgenus: Neja Motsch., 1864 Subgenus: Chlorodium Motsch., 1864 Subgenus: Metallina Motsch., 1850 = Leja Motsch., 1844 Subgenus: Pseudometallina Net., 1920 Subgenus: Phyla Motsch., 1844 = Phayla Motsch., 1844 = Phaula Bed., 1879 = Microcys J.Sahlb., 1907 = Phila auct., incorr. emend. Subgenus: Princidium Motsch., 1864 Subgenus: Paraprincidium Net., 1914 Subgenus: Testedium Motsch., 1864 Subgenus: Actedium Motsch., 1864 Subgenus: Notaphus Dej., 1821 Subgenus: Eupetedromus Net., 1911 = Eupetodromus auct. Subgenus: Notaphemphanes Net., 1920 Subgenus: Notaphocampa Net., 1914 Subgenus: Philochtus Steph., 1829 = Campa Motsch., 1844 Subgenus: Emphanes Motsch., 1850 = Omala Motsch., 1844 Subgenus: Talanes Motsch., 1864 Subgenus: Necpericompsus Net., 1935 Subgenus: Leja Dej., 1821 = Trepanes Motsch., 1864 Subgenus: Trepanedoris Net., 1918 = Trepanoderis Porta, 1923 Subgenus: Semicampa Net., 1910 Subgenus: Diplocampa Bed., 1896 = Campa Bed., 1879 = Paralopha Casey, 1918 Subgenus: Bembidion Latr., 1802 = Lopha Dej., 1821 Subgenus: Nepha Motsch., 1864 = Bembidion auct. Subgenus: Bembidionetolitzkya Strand, 1929 = Daniela Net., 1910 Subgenus: Plataphus Motsch., 1864 = Parataphus Jedl., 1932 Subgenus: Plataphodes Ganglb., 1892 Subgenus: Hirmoplataphus Net., 1943 Subgenus: Trichoplataphus Net., 1914 = Blepharoplataphus Net., 1920 Subgenus: Lymnaeoperyphus Nak., 1963 Subgenus: Limnaeum Steph., 1929 Subgenus: Peryphiolus Jeann., 1941 Subgenus: Euperyphus Jeann., 1941 Subgenus: Ocydromus Clairv., 1806 = Peryphus Steph., 1829 Subgenus: Testediolum Ganglb., 1892 = Peryphidium Tschit., 1895 Subgenus: Pamirium Net., 1920 Subgenus: Synechostictus Motsch., 1864 Subgenus: Pseudolimnaeum Kr., 1888 Tribus: POGONINI Genus: Olegius Komarov, in litt Genus: Cardioderus Dej., 1829 Genus: Pogonus Dej., 1822 Subgenus: Pogonoidius Carret, 1903 = Pogonulus Lutsh., 1926 Subgenus: Pogonus Nic., 1822 = Nogopus Lutsh., 1926 = Raptor Lutsh., 1926 = Calopogonus Lutsh., 1926 Genus: Pogonistes Chaud., 1870 Subgenus: Pogonistes Chaud., 1870 = Eupogonistes Carret, 1903 Subgenus: Syrdenus Chaud., 1870 Genus: Bedeliolus Sem., 1900 Supertribus: PATROBITAE Tribus: PATROBINI Genus: Patrobus Dej., 1821 = Geopatrobus Darl., 1938 Genus: Diplous Motsch., 1850 = Platidius Chaud., 1871 Tribus: DELTOMERINI Genus: Deltomerus Motsch., 1850 = Platynus Dej., 1831 = Cardiomera Chaud., 1846 Genus: Platydiolus Chaud., 1878 = Patroboidea Dyke, 1925 Supertribus: PSYDRITAE Tribus: PSYDRINI Genus: Nomius Laporte, 1834 = Haplochile Lec., 1850 Supertribus: PTEROSTICHITAE Tribus: MORIONINI Genus: Morion Latr., 1810 = Morio auct. Tribus: PTEROSTICHINI Genus: Stomis Clairv., 1806 Subgenus: Stomis Clairv., 1806 Subgenus: Eustomis Sem., 1889 Genus: Abacetus Dej., 1828 Subgenus: Astigis Rambur, 1838 Genus: Poecilus Bon., 1810 Subgenus: Poecilus Bon., 1810 = Feronia Latr., 1817 = Sogines Steph., 1828 = Metapedius Fiori, 1903 = Macropoecilus Lutsh., 1924 = Parapoecilus Jeann., 1941 = Americobius Lutsh., 1914 = Leconteus Lutsh., 1914 = Coelipus Lutsh., 1914 = Pseudosogines Lutsh., 1914 = Glazunovius Lutsh., 1914 = Praveius Lutsh., 1914 Subgenus: Ancholeus Dej., 1828 = Angoleus A.Villa, 1833 = Pseudopedius Seidl., 1887 = Parapedius Tschit., 1898 Subgenus: Derus Motsch., 1850 = Derulus Tschit., 1896 Subgenus: Pseudoderus Seidl., 1887 Subgenus: Lyropedius Seidl., 1887 Genus: Pterostichus Bon., 1810 = Platysma auct. Subgenus: Platysma Bon., 1810 Subgenus: Lyperopherus Motsch., 1850 = Metallophilus Motsch., 1844 Subgenus: Tundraphilus O.Berlov, 1996 Subgenus: Myosodus Fisch., 1823 = Glyptopterus Chaud., 1838 Subgenus: Plectes Fisch., 1822 Subgenus: Natalianoe O.Berlov & Plutenko, 1997 = Paralianoe Ishida, 1958 Subgenus: Galapterus O.Berlov & Plutenko, 1997 Subgenus: Argutor Dej., 1828 = Lagarus Chaud., 1838 = Pseudargutor Casey, 1918 = Eolagarus Tschit., 1899 = Paralagarus Lutsh., 1922 = Pseudolagarus Lutsh., 1922 Subgenus: Pledarus Motsch., 1865 = Badistrinus Motsch., 1865 = Ragadus Motsch., 1865 = Eurythorax Tschit., 1888 = Pseudadelosia Tschit., 1888 = Rhagadulus Tschit., 1897 Subgenus: Pedius Motsch., 1850 Subgenus: Adelosia Steph., 1835 = Agonodemus Chaud., 1837 Subgenus: Melanius Bon., 1810 = Pseudomaseus Chaud., 1838 = Micromaseus Desbroch., 1906 Subgenus: Phonias Goz., 1886 = Argutor auct. = Biphonias Jeanne, 1988 Subgenus: Cryobiopterus O.Berlov, 1997 Subgenus: Cryobius Chaud., 1838 = Pseudocryobius Motsch., 1850 = Orites Schaum, 1860 Subgenus: Haptoderus Chaud., 1838 = Pseudorthomus Chaud., 1838 Subgenus: Oreoplatysma Jacobs., 1906 = Agonodemus Rtt., 1896 Subgenus: Asioplatysma Kryzh., 1968 = Pseudoderus auct. part. Subgenus: Haplomaseus Rtt., 1896 Subgenus: Eurymelanius Rtt., 1896 Subgenus: Lyrothorax Chaud., 1838 Subgenus: Omaseus Dej., 1821 = Lyperus Chaud., 1838 = Lyperosomus Motsch., 1850 Subgenus: Steropus Dej., 1821 = Steroderus Motsch., 1850 Subgenus: Eosteropus Tschit., 1902 Subgenus: Lenapterus O.Berlov, 1996 = Steroperis Shil., 1995 Subgenus: Bothriopterus Chaud., 1838 Subgenus: Morphnosoma Lutsh., 1915 = Omaseus auct. = Omaseidius Jeann., 1942 = Euferonia Casey, 1918 Subgenus: Feronidius Jeann., 1941 = Pterostichus Schaum, 1860 Subgenus: Cophosus Dej., 1828 Subgenus: Falsargutor Kryzh., 1984 Subgenus: Petrophilus Chaud., 1838 = Euryperus Motsch., 1850 = Feroperis Lafer, 1980 Subgenus: Oreolyperus Tschit., 1901 = Boreobia Tschit., 1896 Subgenus: Stereocerus Kirby, 1837 Subgenus: Bryobius Chaud., 1838 = Psychobius Chaud., 1838 Subgenus: Oreophilus Chaud., 1838 Subgenus: Cheporus Latr., 1825 Subgenus: Calopterus Chaud., 1838 = Dasalus Motsch., 1850 = Aello Goz., 1882 Subgenus: Agastillus Rtt., 1892 Genus: Aphaonus Rtt., 1887 Subgenus: Aphaonus Rtt., 1887 Subgenus: Scaritomorphus Kurn., 1962 Genus: Abax Bon., 1810 Genus: Molops Bon., 1810 Tribus: SPHODRINI : Calathi Genus: Calathus Bon., 1810 Subgenus: Calathus Bon., 1810 Subgenus: Neocalathus Ball & Negre, 1972 Subgenus: Dolichus Bon., 1809 Subgenus: Lindrothius Kurn., 1961 Genus: Thermoscelis Putz., 1873 : Sphodri Genus: Pseudotaphoxenus Schauf., 1865 = Rhopalomelus Motsch., 1864 Genus: Stenolepta Sem., 1889 Genus: Taphoxenus Motsch., 1864 Subgenus: Taphoxenus Motsch., 1864 Subgenus: Lychnifugus Motsch., 1864 Genus: Reflexisphodrus Casale, 1988 Genus: Sphodrus Clairv., 1806 Genus: Eremosphodrus Sem., 1908 Subgenus: Erempsphodrus Sem., 1908 Subgenus: Rugisphodrus Casale, 1988 Genus: Laemostenus Bon., 1810 = Laemosthenes Schauf., 1865 Subgenus: Laemostenus Bon., 1810 Subgenus: Antisphodroides Casale, 1988 Subgenus: Stenosphodrus Casale, 1988 Subgenus: Microsphodrus Casale, 1988 Subgenus: Antisphodrus Schauf., 1865 Subgenus: Pristonychus Dej., 1828 = Cryptoxenus Motsch., 1864 Tribus: PLATYNINI = AGONINI = ANCHOMENINI Genus: Sericoda Kirby, 1837 = Rhytiderus Chaud., 1844 = Tanystoma Rtt., 1907 = Agonodromius Rtt., 1908 Genus: Agonum Bon., 1810 Subgenus: Agonum Bon., 1810 Subgenus: Liebherrius Shil. Subgenus: Europhilus Chaud., 1859 Subgenus: Platynomicrus Casey, 1920 ? Agonocyrthes Motsch., 1864 Genus: Platynus Bon., 1810 Subgenus: Platynus Bon., 1810 = Limodromus Motsch., 1864 Subgenus: Batenus Motsch., 1864 = Platynidius Casey, 1920 = Pseudoplatynus Habu, 1973 Genus: Paranchus Lindr., 1974 Genus: Oxypselaphus Chaud., 1843 = Anchus Lec., 1854 Genus: Anchomenus Bon., 1810 = Clibinarius Goz., 1882 = Idiochroma Bed., 1902 Genus: Anchodemus Motsch., 1864 Genus: Chlaeniomimus Sem., 1889 Genus: Anchagonum Kryzh., 1995 Genus: Nipponanchus Habu, 1978 = Pseudanchus Casey, 1920 Genus: Eucolpodes Jeann., 1948 Subgenus: Eucolpodes Jeann., 1948 Subgenus: Glaucagonum Habu, 1978 Genus: Gyrochaetostylus Habu, 1978 Genus: Metacolpodes Jeann., 1948 Genus: Dicranoncus Chaud., 1878 Genus: Olisthopus Dej., 1828 = Odontonyx auct. Genus: Atranus Lec., 1848 Subtribus: SYNUCHINA Genus: Synuchus Gyll., 1810 = Taphria Dej., 1828 Subgenus: Synuchus Gyll., 1810 Subgenus: Pristodactyla Dej., 1828 = Paracalathus Jedl., 1940 = Semenovia Jedl., 1953 Subgenus: Crepidactyla Motsch., 1861 = Crepidodactyla Jacobs., 1907 = Fuerthius Jedl., 1953 Genus: Platyderus Steph., 1828 = Lissotarsus Chaud., 1838 Subtribus: PRISTOSIINA Genus: Pristosia Motsch., 1864 = Eucalathus Bat., 1883 = Kanoldia Jedl., 1931 = Laemostenopsis Jedl., 1931 Tribus: AMARINI Genus: Amara Bon., 1810 Subgenus: Zezea Csiki, 1929 = Triaena Lec., 1848 = Pseudotriaena Minsk & Hatch, 1939 Subgenus: Amara Bon., 1810 = Pseudoamara Bal., 1934 Subgenus: Celia Zimm., 1832 = Acrodon Zimm., 1831 = Isopleurus Kirby, 1837 = Amarocelia Motsch., 1862 = Bradycelia Lafer, 1989 Subgenus: Reductocelia Lafer, 1989 Subgenus: Paracelia Bed., 1899 Subgenus: Oreoamara Lutsh., 1927 Subgenus: Iranoleiridis Hieke, 1978 Subgenus: Bradytulus Tschit., 1894 Subgenus: Bradytus Steph., 1828 = Pseudobradytus Csiki, 1908 = Pseudocelia Lutsh., 1935 Subgenus: Heterodema Tschit., 1894 Subgenus: Ammoxena Tschit., 1894 Subgenus: Eoleirides Tschit., 1898 Subgenus: Percosia Zimm., 1832 Subgenus: Embrikiella Lutsh., 1935 Subgenus: Parapercosia Tschit., 1898 Subgenus: Pseudoleirides Kryzh., 1968 = Tadzhikamara Kryzh., 1968 Subgenus: Amathitis Zimm., 1832 Subgenus: Zabroscelis Putz., 1866 Subgenus: Leiramara Hieke, 1988 Genus: Harpalodema Rtt., 1888 Genus: Phanerodonta Tschit., 1894 Genus: Cribramara Kryzh., 1964 Genus: Curtonotus Steph., 1828 = Cyrtonotus auct. = Leirus Zimm., 1832 Subgenus: Curtonotus Steph., 1828 = Leirodema Tschit., 1894 = Feronalius Casey, 1918 = Paracurtonotus Habu, 1942 = Paracyrtonotus Bal., 1943 Subgenus: Allobradytus Khnz., 1975 Subgenus: Ammoleirus Tschit., 1899 Subgenus: Microleirus Kryzh., 1974 Genus: Polysitamara Kryzh., 1968 Genus: Zabrus Clairv., 1806 = Pelobatus Fisch., 1817 Subgenus: Zabrus Clairv., 1806 Subgenus: Pelor Bon., 1810 = Pelorosomus Motsch., 1850 Subgenus: Eutroctes Zimm., 1831 Supertribus: HARPALITAE Tribus: HARPALINI Subtribus: ANISODACTYLINA Genus: Anisodactylus Dej., 1829 Subgenus: Anisodactylus Dej., 1829 Subgenus: Pseudanisodactylus Noonan, 1973 = Morphogynandrus Garrett, 1905 Subgenus: Pseudodichirus Lutsh., 1921 Subgenus: Hexatrichus Tschit., 1898 Genus: Gynandromorphus Dej., 1829 Genus: Diachromus Er., 1837 Subtribus: STENOLOPHINA Genus: Bradycellus Er., 1837 = Tetraplatypus Tschit., 1897 Subgenus: Bradycellus Er., 1837 Subgenus: Tachycellus Mor., 1862 Subgenus: Stenocellus Casey, 1914 Genus: Dicheirotrichus Jacq., 1857 Subgenus: Dicheirotrichus Jacq., 1857 Subgenus: Cardiostenus Rtt., 1900 Subgenus: Oreoxenus Tschit., 1899 Subgenus: Trichocellus Ganglb., 1892 Genus: Lioholus Tschit., 1897 Genus: Stenolophus Steph., 1828 Subgenus: Stenolophus Steph., 1828 Subgenus: Egadroma Motsch., 1855 Genus: Idiomelas Tschit., 1900 = Hemiaulax Bat., 1892 Subgenus: Idiomelas Tschit., 1900 Genus: Loxoncus Schm.-Goeb., 1846 = Anoplogenus Chaud., 1852 = Megrammus Motsch., 1857 Genus: Acupalpus Latr., 1829 = Manicellus Motsch., 1864 Subgenus: Ancylostria Schaub., 1930 Subgenus: Acupalpus Latr., 1829 Subgenus: Setacupalpus Habu, 1973 Subgenus: Palcuapus Habu, 1973 Genus: Anthracus Motsch., 1850 = Balius Schiodte, 1861 Subtribus: HARPALINA Genus: Daptus Fisch., 1824 Genus: Trichotichnus Mor., 1863 = Iridessus Bat., 1883 = Asmerinx Tschit., 1898 Genus: Nipponoharpalus Habu, 1973 Genus: Parophonus Ganglb., 1892 Subgenus: Parophonus Ganglb., 1892 Subgenus: Tachyophonus Tschit., 1900 Subgenus: Ophonomimus Schaub., 1923 Genus: Harpalobrachys Tschit., 1899 Genus: Harpalus Latr., 1802 = Haploharpalus Schaub., 1926 = Euharpalops Casey, 1924 = Harpalobius Rtt., 1900 = Harpalellus Lindr., 1968 = Conicus Motsch., 1844 = Pheuginus Motsch., 1844 = Hypsinephus Bat., 1878 = Rapahlus Lutsh., 1922 = Artabas Goz., 1882 = Harpalophonus Ganglb., 1892 = Epiharpalus Rtt., 1900 Genus: Neophygas Noonan, 1976 = Phygas Motsch., 1848 Genus: Microderes Fald., 1835 Subgenus: Microderes Fald., 1835 = Bioderus Motsch., 1852 Subgenus: Neopangus Tschit., 1898 Subgenus: Microharpalus Tschit., 1898 Genus: Pangus Dej., 1821 Genus: Acinopus Dej., 1821 Subgenus: Acinopus Dej., 1821 Subgenus: Haplacinopus Sem., 1899 Subgenus: Osimus Motsch., 1850 Subgenus: Oedematicus Bed., 1897 Genus: Bleusea Bed., 1896 Genus: Ophonus Dej., 1821 Subgenus: Brachyophonus Sciaky, 1987 = Graecophonus Schaub. Subgenus: Metophonus Bed., 1897 Subgenus: Hesperophonus Ant., 1959 Subgenus: Ophonus Dej., 1821 Subgenus: Macrophonus Tschit., 1901 Genus: Penthophonus Rtt., 1900 Subtribus: DITOMINA Genus: Penthus Chaud., 1843 Genus: Liochirus Tschit., 1897 Genus: Graniger Motsch., 1864 = Carterophonus Ganglb., 1890 Genus: Eucarterus Rtt., 1900 Genus: Oedesis Motsch., 1850 = Eriotomus Pioch., 1873 = Eriocypas Tschit., 1901 Genus: Carterus Dej., 1829 = Distomus Steph., 1827 = Odogenius Sol., 1835 Subgenus: Carterus Dej., 1829 Subgenus: Pristocarterus Wrase, 1994 Genus: Tschitscherinellus Csiki, 1906 = Ditomus auct. non Bon., 1810 = Gonoxyaristus Stichel, 1923 Genus: Ditomus Bon., 1810 = Aristus Latr., 1816 = Sabienus Goz., 1882 = Euditomus Acloque, 1896 Genus: Dixus Billb., 1820 Genus: Proditomus Schaub., 1934 Genus: Eocarterus Stichel, 1923 = Apterocarterus Stichel, 1923 Genus: Bronislavia Sem., 1891 Genus: Carenochyrus Sols., 1874 Genus: Machozethus Chaud., 1850 = Harpactes Men., 1849 = Dioctes Men., 1849 Genus: Chilotomus Chaud., 1842 = Chlochilus Motsch., 1850 Subtribus: AMBLYSTOMINA Genus: Amblystomus Er., 1837 = Hispalis Rambur, 1838 = Artizoum Gistl, 1857 = Megaristerus Nietner, 1858 = Notophilus Blackburn, 1888 = Thenarotidius Sloane, 1893 = Psilonothus Sloane, 1899 = Entomorrhinus Jeann., 1948 Supertribus: PERIGONITAE Tribus: PERIGONINI Genus: Perigona Cast., 1835 = Extromus Peringuey, 1896 ? Nestra Motsch., 1851 ? Spathinus Nietner, 1858 Subgenus: Trechicus Lec., 1853 Supertribus: PANAGAEITAE Tribus: PANAGAEINI Genus: Panagaeus Latr., 1802 Genus: Tinoderus Chaud., 1878 Genus: Peronomerus Schaum, 1853 Supertribus: CALLISTITAE Tribus: CALLISTINI Genus: Callistus Bon., 1809 Genus: Eochlaenius Sem., 1912 Genus: Epomis Bon., 1810 Genus: Dinodes Bon., 1810 = Glyptoderus auct. = Paradinodes Apf., 1904 Genus: Chlaenius Bon., 1810 Subgenus: Lissauchenius MacLeay, 1825 = Ilaenchus Lutsh., 1933 Subgenus: Stenochlaenius Rtt., 1908 = Turanochlaenius Lutsh., 1933 Subgenus: Trichochlaenius Seidl., 1887 Subgenus: Litochlaenius Kryzh., 1976 = Hemichlaenius Lutsh., 1933 Subgenus: Chlaenites Motsch., 1860 Subgenus: Chlaenius Bon., 1810 = Chaelinus Lutsh., 1933 = Chinelaus Lutsh., 1933 = Laenichus Lutsh., 1933 Subgenus: Chlaeniellus Rtt., 1908 Subgenus: Agostenus Stev., 1829 = Agostenops Lutsh., 1933 Subgenus: Pelasmus Stev., 1829 Tribus: OODINI Genus: Oodes Bon., 1810 Subgenus: Oodes Bon., 1810 Subgenus: Lachnocrepis Lec., 1853 Tribus: LICININI = BADISTRIINI Subtribus: SUBMERINA Genus: Diplocheila Brulle, 1834 = Diplochila auct. = Rembus Dej., 1826 = Rhembus auct. Subgenus: Diplocheila Brulle, 1834 Subgenus: Submera Habu, 1956 Subtribus: LICININA Genus: Licinus Latr., 1802 = Scales Fisch., 1817 Subgenus: Licinus Latr., 1802 Subgenus: Neorescius Heyd. , Rtt. & Weise, 1906 = Orescius Bed., 1879 Subgenus: Tricholicinus Popp., 1912 Genus: Derostichus Motsch., 1859 Genus: Colpostoma Sem., 1889 Genus: Badister Clairv., 1806 = Amblychus Gyll., 1810 Subgenus: Badister Clairv., 1802 Subgenus: Trimorphus Steph., 1828 Subgenus: Baudia Rag., 1884 Supertribus: MASOREITAE Tribus: MASOREINI Genus: Masoreus Dej., 1821 Genus: Mnuphorus Chaud., 1873 Subgenus: 'sellatus' = Mnuphorus Chaud., 1873 Subgenus: 'callistoides' = Hypercosmeton Rtt., 1889 Tribus: TETRAGONODERINI Genus: Tetragonoderus Dej., 1829 = Peronoscelis Chaud., 1876 Tribus: CORSYRINI Genus: Corsyra Dej., 1825 Genus: Discoptera Sem., 1889 Supertribus: PENTAGONITAE Tribus: PENTAGONINI Genus: Pentagonica Schm.-Goeb., 1846 = Rhombodera Reiche, 1842 = Didetus Lec., 1853 Supertribus: ODACANTHITAE Tribus: ODACANTHINI Genus: Odacantha Payk., 1798 = Colliuris auct. = Odontacantha Agass., 1847 = Dictya Chaud., 1870 Supertribus: LEBIITAE Tribus: LEBIINI Subtribus: LEBIINA Genus: Rhopalostyla Chaud., 1850 = Taenioptilon Motsch., 1859 Genus: Lionedya Chaud., 1870 Genus: Lachnolebia Maindr., 1905 Genus: Lebia Latr., 1802 Subgenus: Lamprias Bon., 1809 = Echimuthus Leach, 1815 = Omalomorpha Motsch., 1844 = Homalops Motsch., 1850 = Lebida Motsch., 1862 Subgenus: Lebia Latr., 1802 Subgenus: Poecilothais Maindr., 1905 Subgenus: Glyciolebia Kryzh., 1987 Subgenus: Setolebia Jedl., 1941 Genus: Daer Sem., 1929 Subtribus: CALLIDINA = Neotarus Rtt., 1884 Genus: Lebidia Mor., 1863 Genus: Parena Motsch., 1859 = Crossoglossa Chaud., 1872 Genus: Glycia Chaud., 1842 Subtribus: PLATYTARINA Genus: Cymindoidea Cast., 1832 Subgenus: Platytarus Fairm., 1850 Subtribus: DEMETRIINA Genus: Demetrias Bon., 1810 Subgenus: Demetrias Bon., 1810 Subgenus: Aetophorus Schm.-Goeb., 1846 Subtribus: DROMIINA Genus: Dromius Bon., 1810 Subgenus: Dromius Bon., 1810 Subgenus: Lebidromius Jedl., 1931 Genus: Paradromius Fowl., 1886 Subgenus: Paradromius Fowl., 1886 Subgenus: Manodromius Rtt., 1905 Genus: Philorhizus Hope, 1838 Subgenus: Calodromius Rtt., 1905 Subgenus: Philorhizus Hope, 1838 Genus: Syntomus Hope, 1838 = Metabletus Schm.-Goeb., 1846 Genus: Metadromius Bed., 1886 Genus: Charopterus Motsch., 1839 Genus: Microlestes Schm.-Goeb., 1846 = Blechrus Motsch., 1847 = Bomius Lec., 1851 = Dromius sensu Sloane, 1898 Genus: Lionychus Wissm., 1846 Genus: Apristus Chaud., 1846 = Crephnos Baudi, 1864 Genus: Xanthomelina Khnz., 1964 Genus: Microdaccus Schaum, 1864 Genus: Somotrichus Seidl., 1887 Genus: Plochionus Latr. & Dej., 1824 Subtribus: CYMINDINA Genus: Cymindis Latr., 1806 Subgenus: Baicalotarus Emetz, 1974 Subgenus: Chaetotarus Rtt., 1903 Subgenus: Cymindis Latr., 1806 = Tarus Clairv., 1806 = Anomoeus Fisch., 1822 = Arrhostus Motsch., 1864 = Psammastes Motsch., 1864 Subgenus: Eremocymindis Emetz, 1974 = Psammoxenus Chaud., 1873 Subgenus: Iscariotes Reiche, 1855 Subgenus: Neopsammoxenus Emetz, 1973 Subgenus: Paracymindis Jedl., 1968 = Mastus Motsch., 1864 = Assadeva Mandl, 1973 = Berus Motsch., 1864 Subgenus: Pseudomastus Emetz, 1972 Subgenus: Menas Motsch., 1864 Subgenus: Neomenas Emetz, 1974 Subgenus: Pseudocymindis Habu, 1967 Subgenus: Tarsostinus Motsch., 1864 Subgenus: Tarulus Bed., 1906 Genus: Petrimagnia Kryzh. & Mich., 1971 Genus: Agatus Motsch., 1844 Genus: Merizomena Chaud., 1872 Genus: Trichis Klug, 1831 Genus: Anomotarus Chaud., 1875 Tribus: ANTHIINI Genus: Anthia Web., 1801 Subgenus: Pachymorpha Hope, 1838 Tribus: DRYPTINI Genus: Drypta Latr., 1796 Tribus: ZUPHIINI Genus: Polistichus Bon., 1809 Genus: Zuphium Latr., 1806 = Zophium Gistl, 1838 = Zoyphium Motsch., 1850 Subgenus: Zuphium Latr., 1806 Subgenus: Parazuphium Jeann., 1942 Subgenus: Neozuphium Hurka, 1982 Subfamilia: BRACHININAE Tribus: BRACHININI Genus: Aptinus Bon., 1810 Genus: Brachinus Web., 1801 = Brachynus auct. = Brachynidius Rtt., 1919 = Brachynolomus Rtt., 1919 = Cnecostolus Rtt., 1919 Genus: Mastax Fisch., 1827 Genus: Pheropsophus Sol., 1833 Subfamilia: PAUSSINAE Tribus: PAUSSINI Genus: Paussus L., 1775