| Selected Publications: |
- Orlova V.F., Bakharev V.A. & Borkin L.J. 1977. Karyotypes of some brown frogs of Eurasia and a taxonomic analysis of karyotypes of the group. Proceedings of the Zoological Institute, USSR Academy of Sciences, Leningrad, 74: 84–103 (in Russian, with English summary).
- Borkin L.J. & Darevsky I.S. (1980). Reticulate (hybridogenous) speciation in vertebrates. Zhurnalf Obshchei Biologii, Moscow, 41 (4): 485–506 (in Russian, with English abstract).
- Borkin L. J.1984. The European–Far Eastern disjunctions in distribution of amphibians: a new analysis of the problem. In: L.J. Borkin (ed.). Ecology and Faunistics of Amphibians and Reptiles of the USSR and Adjacent Counties. Proceedings of the Zoological Institute, USSR Academy of Sciences, Leningrad, 124: 55–88 (in Russian, with English summary).
- Matsui M., Seto T., Kohsaka Y., Borkin L.J. 1985. Bearing of chromosome C-banding patterns on the classification of Eurasian toads of the Bufo bufo complex. Amphibia-Reptilia, Leiden, 6 (1): 23–33.
- Borkin, L. J., A. E. Vinogradov, J. M. Rosanov, and I. A. Caune. 1987. Hemiclonal inheritance in the hybridogenetic complex Rana esculenta: evidence from DNA flow cytometry. Doklady Akademii Nauk SSSR, 295 (5):1261–1264 (in Russian).
- Ananjeva N.B., Borkin L.J., Darevsky I.S., Orlov N.L. 1988. Dictionary of Animal Names in Five Languages. Amphibians and Reptiles. Latin, Russian, English, German, French. 12 126 names. Moscow: Russky Yazyk Publishers, 556 p.
- Borkin L.J., Kuzmin S.L. 1988. Amphibians of Mongolia: species accounts. In: Vorobyeva E.I., Darevsky I.S. (eds). Amphibians and Reptiles of MNR. General Problems. Amphibians. Moscow: Nauka, p. 30–197 (in Russian).
- Vinogradov A.E., Borkin L.J., Günther R., Rosanov J.M. 1990. Genome elimination in diploid and triploid Rana esculenta males: cytological evidence from DNA flow cytometry. Genome, 33 (5): 619–627. https://doi.org/10.1139/g90-092
- Caune I.A., Borkin L.J. 1993. A new kind of unisexual-bisexual population systems in European green frogs (Rana esculenta complex). In: Panov E.N. & Tomkovich P.S. (eds.). Hybridization and Species Problem in Vertebrates [Hibridisazija i problema vida u pozvonochnykh]. Moscow, Archives of Zoological Museum, Moscow State University, vol. 30, p. 34–52 (in Russian, with English summary).
- Green D.M., Borkin L.J. 1993. Evolutionary relationships of eastern Palearctic brown frogs, genus Rana: paraphyly of the 24-chromosome species group and the significance of chromosome number change. Zoological Journal of the Linnean Society, 109 (1): 1–25. https://doi.org/10.1111/j.1096-3642.1993.tb01256.x
- Miura I., Nishioka M., Borkin L.J. & Wu Z. 1995. The origin of the brown frogs with 2n=24 chromosomes. Experientia, Basel, 51 (2): 179–188. https://doi.org/10.1007/BF01929366
- Ananjeva N.B., Borkin L.J., Darevsky I.S., Orlov N.L. 1998. Amphibians and Reptiles. AFB, Moscow, 574 p. (in the series: Encyclopedia of Russia’s Nature; in Russian).
- Borkin L.J. 1999. Distribution of amphibians in North Africa, Europe, Western Asia, and the former Soviet Union. In: Duellman W.E. (ed.). Patterns of Distribution of Amphibians: a Global Perspective. Baltimore and London: The Johns Hopkins University Press, 329–420.
- Borkin L.J., Eremchenko V.K., Helfenberger N., Panfilov A.M., Rosanov J.M. 2001. On the distribution of diploid, triploid, and tetraploid green toads (Bufo viridis complex) in south-eastern Kazakhstan. Russian Journal of Herpetology, 8 (1): 45–53, and 8(3): 246 (errata).
- Litvinchuk S.N. & Borkin L.J. 2003. Variation in number of trunk vertebrae and in count of costal grooves in salamanders of the family Hynobiidae. Contributions to Zoology, The Hague, 72 (4): 195–209. doi: https://doi.org/10.1163/18759866-0720400
- Babik W., Branicki W., Sandera M., Litvinchuk S., Borkin L.J., Irwin J.T., Rafinski J. 2004. Mitochondrial phylogeography of the moor frog, Rana arvalis. Molecular Ecology, 13: 1469–1480.
- Flyaks N.L., Borkin L.J. 2004. Morphological abnormalities and heavy metal concentrations in anurans of contaminated areas, eastern Ukraine. Applied Herpetology, Leiden, 1 (3–4): 229–264.
- Litvinchuk S.N., Borkin L.J. & Rosanov J.M. 2004. Intraspecific and interspecific genome size variation in hynobiid salamanders of Russia and Kazakhstan: determination by flow cytometry. Asiatic Herpetological Research, Berkeley, 10: 282–294.
- Borkin, L.J., S.N. Litvinchuk, J.M. Rosanov & D.V. Skorinov (2004): On cryptic species (from the example of amphibians). Zoologicheskii Zhurnal, Moscow 83/8: 936–960 [in Russian; full English version in Entomological Review, Moscow 84/Suppl. 1: S75–S98].
- Borkin L.J., Korshunov A.V., Lada G.A., Litvinchuk S.N., Rosanov J.M., Shabanov D.A., Zinenko A.I. 2004. Mass occurrence of polyploid green frogs (Rana esculenta complex) in eastern Ukraine. Russian Journal of Herpetology, 11(3): 203–222.
- Borkin L.J. 2009. Carl Linnaeus (1707–1778) as a zoologist. Proceedings of the Zoological Institute, Russian Academy of Sciences, St. Petersburg, Suppl. 1: 9–78 (Alimov A.F. & Stepan’yants S.D., eds. Species and Speciation. The analysis of new views and trends).
- Litvinchuk S. N., Borkin L. J. 2009. Evolution, systematics and distribution of crested newts (Triturus cristatus complex) in Russia and adjacent countries. St. Petersburg: Evropeisky Dom Press, 592 p., 24 p. color. ill.
- Borkin L.J. & Litvinchuk S.N. (2010). Species and speciation: a molecular-genetic evaluation (genetic distances). In: Kolchinsky E.I. (ed.). Charles Darwin and Contemporary Biology. St. Petersburg: Nestor-Istoria, p. 236–250 (in Russian, with English summary).
- Hasumi M., Borkin L.J. 2012. Age and body size of Salamandrella keyserlingii (Caudata: Hynobiidae): a difference in altitudes, latitudes, and temperatures. Organisms Diversity & Evolution, 12 (2): 167–181.
- Plötner J., Baier F., Akın C., Mazepa G., Schreiber R., Beerli P., Litvinchuk S.N., Bilgin C.C., Borkin L., Uzzell T. 2012. Genetic data reveal that water frogs of Cyprus (genus Pelophylax) are an endemic species of Messinian origin. Zoosystematics and Evolution, 88 (2): 261–283.
- Borkin L.J., Bezman O.S., Litvinchuk S.N. 2012. The evaluation of occurrence of anomalies in natural populations (on example of amphibians). Proceedings of the Zoological Institute, Russian Academy of Sciences, St. Petersburg, 316 (4): 324–343 (in Russian, with English summary). http://www.zin.ru/journals/trudyzin/doc/vol_316_4/TZ_316_4_Borkin.pdf
- Litvinchuk S.N., Crottini A., Federici S., De Pous P., Donaire D., Andreone F., Kalezić M.L., Džukić G., Lada G.A., Borkin L.J., Rosanov J.M. 2013. Phylogeographic patterns of genetic diversity in the common spadefoot toad, Pelobates fuscus (Anura: Pelobatidae), reveals evolutionary history, postglacial range expansion and secondary contact. Organisms Diversity and Evolution, 13 (3): 433–451.
- Dedukh D., Mazepa G., Shabanov D., Rosanov J., Litvinchuk S., Borkin L., Vences M., Hauswaldt J.S., Steinfartz S., Rupp O., Goesmann A., Künzel S., Orozco-terWengel P., Vieites D.R., Nieto-Roman S., Haas S., Laugsch C., Gehara M., Bruchmann S., Pabijan M., Ludewig A.-K., Rudert D., Angelini C., Borkin L.J., Crochet P.-A., Crottini A., Dubois A., Ficetola G.F., Galán P., Geniez Ph., Hachtel M., Jovanovic O., Litvinchuk S.N., Lymberakis P., Ohler A., Smirnov N.A. 2013. Radically different phylogeographies and patterns of genetic variation in two European brown frogs, genus Rana. Molecular Phylogenetics and Evolution, 68 (3): 657–670.
- Borkin L.J., Litvinchuk S.N. 2013. Hybridization, speciation, and systematics of animals. Proceedings of the Zoological Institute, Russian Academy of Sciences, St. Petersburg, Suppl. 2: 83–139 (Alimov A.F. & Stepan’yants S.D., eds. Contemporary Problems of Biological Systematics). http://www.zin.ru/journals/trudyzin/doc/vol_317_s1/TZ_317_1_Supplement_Borkin.pdf
- Borkin L.J., Litvinchuk S.N. 2013. Amphibians of the Palearctic: taxonomic composition. Proceedings of the Zoological Institute, Russian Academy of Sciences, St. Petersburg, 317 (4): 494–541 (in Russian, with English summary). http://www.zin.ru/journals/trudyzin/doc/vol_317_4/TZ_317_4_Borkin_Litvinchuk.pdf
- Gollmann B., Borkin L., Grossenbacher K., Weddeling K. 2014. Rana temporaria Linnaeus 1758 – Grasfrosch. In: Grossenbacher K. (Hrsg.). Handbuch der Reptilien und Amphibien Europas. Band 5/III A: Froschlurche (Anura) III A (Ranidae I). Wiebelsheim: AULA-Verlag, S. 305–437.
- Borkin L.J. 2014. Morphological abnormalities in natural populations of amphibians: what do we study and how do we measure? In: Vershinin V.A., Dubois A., Henle K. & Puky M. (eds.). Anomalies and Patologies in Amphibians and Reptiles: Methodology, Evolutionary Importance, Possibilities of Environmental Health Assessment. Yekaterinburg: Yekaterinburg University Press, p. 25–36 (in Russian, with English summary).
- Borkin L.J. 2014. Izvara, N.K. Roerich, the Himalaya. St. Petersburg: Evropeisky Dom, 254 p. (in Russian)
- Litvinchuk S.N., Schepina N.A., Munkhbaatar M., Munkhbayar Kh., Borkin L.J., Kazakov V.I., Skorinov D.V. 2014. Distribution and conservation status of the Far Eastern tree frog, Hyla japonica Günther, 1859 in Mongolia and Transbaikalia (Russia). Russian Journal of Herpetology, 21 (4): 303–314.
- Borkin L.J., Litvinchuk S.N. 2014. Zoogeography of Northern Hemisphere and amphibians: the Palearctic and the Nearctic realms or the Holarctic Realm? Proceedings of the Zoological Institute, Russian Academy of Sciences, St. Petersburg, 318 (4): 433–485 (in Russian, with English summary). http://www.zin.ru/journals/trudyzin/doc/vol_318_4/TZ_318_4_Borkin.pdf
- Borkin L.J., Gannibal B.K., Golubev A.V. 2014. On the pathways of Peter Simon Pallas (journeys through western Kazakhstan). St. Petersburg – Uralsk: Eurasian Union of Scholars, 310 p. (in Russian).
- Litvinchuk S.N., Borkin L.J., Litvinchuk Y.S., Rosanov J.M. 2015. Distribution and population systems of green frogs (Pelophylax esculentus complex) in Kaliningrad oblast’, Russia (Baltic Sea region). Russian Journal of Herpetology, 22 (3): 188–196.
- Wagner P., Tiutenko A., Mazepa G., Borkin L.J., Simonov E. 2016. Alai! Alai! – a new species of the Gloydius halys (Pallas, 1776) complex (Viperidae, Crotalinae), including a brief review of the complex. Amphibia-Reptilia, Leiden, 37 (1): 15–31. doi: https://doi.org/10.1163/15685381-00003026
- Biriuk O.V., Shabanov D.A., Korshunov A.V., Borkin L.J., Lada G.A., Pasynkova R.A., Rosanov J.M., Litvinchuk S.N. 2016. Gamete production patterns and mating systems in water frogs of the hybridogenetic Pelophylax esculentus complex in north-eastern Ukraine. Journal of Zoological Systematics and Evolutionary Research, 54 (3): 215–225. doi: 10.1111/jzs.12132
- Borkin L.J. 2016. In defense of Ivan Efremov, a paleontologist and science fiction writer (1974). Studies in the History of Biology, St. Petersburg, 8 (1): 94–120 (in Russian, with English summary).
- Borkin L.J. & Litvinchuk S.N. (2016). On amphibian fauna of Tibet (a preliminary analysis). Vestnik of St. Petersburg University, series 3, biology, 3: 25–30 (in Russian, with English summary).
- Litvinchuk S. N., Borkin L. J., Skorinov D. V., Pasynkova R.A. & Rosanov J.M. 2016. Natural polyploidy in amphibians. Vestnik of St. Petersburg University, series 3, biology, 3: 77–86. https://doi.org/10.21638/11701/spbu03.2016.314 (in Russian, with English abstract).
- Borkin L.J., Goncharov A.I., Litvinchuk S.N. 2016. The Egyptian toad, Sclerophrys regularis (Reuss, 1833) at Sharm El-Sheikh, with comments on amphibians of the Sinai Peninsula. Russian Journal of Herpetology, 23 (4): 283–292.
- Borkin L.J. 2016. Professor Leszek Berger and his influence on green frog studies in Russia, with my recollections. In: Szajdak L.W., Śmiełowski J. (eds). Impact of Professor Leszek Berger’s Discoveries on the Develoment of Biological Sciences. Poznań: Instytut Środowiska Rolniczego i Leśnogo Polskiej Akademii Nauk, p. 51–88.
- Litvinchuk S.N., Borkin L.J., Mazepa G., Skorinov D.V., Melnikov D.A. & Rosanov J.M. 2017. Distribution of amphibians in the Western Himalaya (India). In: L.J. Borkin (ed.). Russian Himalayan Research: past, present, future. St. Petersburg: Evropeisky Dom, p. 188–194 (in Russian).
- Litvinchuk S.N., Skorinov D.V., Mazepa Glib O., Borkin L.J. 2018. Distribution of Bufotes latastii (Boulenger, 1882), endemic to the Western Himalaya. Alytes, Paris, 36 (1–4): 314–327.
- Faizulin A.I., Svinin A.O., Ruchin A.B., Skorinov D.V., Borkin L.J., Rosanov J.M., Kuzovenko A.E. & Litvinchuk S.N. (2018). Distribution and contact zone of two forms of greens toads of the Bufotes viridis complex (Anura, Amphibia), different in genome size, in the Volga Region. Current Studies in Herpetology, Saratov, 18 (1/2): 35–45. https://doi.org/10.18500/1814-6090-2018-18-1-2-35-45 (in Russian, with English summary)
- Borkin L.J., Litvinchuk S.N., Melnikov D.A., Skorinov D.V. 2018. Altitudinal distribution of skinks of the genus Asymblepharus in the Western Himalaya, India (Reptilia: Sauria: Scincidae). In: Hartmann M., Barclay M.V.L., Weipert J. (Hrsg.). Biodiversität und Naturausstattung im Himalaya VI. Erfurt: Verein der Freunde & Förderer des Naturkundemuseums Erfurt e.V., p. 163–167.
- Litvinchuk S.N., Borkin L.J., Mazepa G. & Rosanov J.M. 2018. Genome size and distribution of diploid and polyploid green toads of the genus Bufotes in Uzbekistan and Turkmenistan. In: Herpetological and Ornithological research: current aspects. Moscow: KMK Scientifi c Press. 2018. P. 88–101 (in Russian, with English summary).
- Litvinchuk S.N., Melnikov D.A., Borkin L.J., Hofmann S. 2019. Rediscovery of the high altitude lazy toad, Scutiger occidentalis Dubois, 1978, in India. Russian Journal of Herpetology, 26 (1): 17–22. https://doi.org/10.30906/1026-2296-2019-26-1-17-22
- Litvinchuk S.N., Skorinov D.V., Pasynkova R.A., Kidov A.A., Matushkina K.A., Borkin L.J., Rosanov J.M. 2019. Polyploid speciation in Asian green toads of the genus Bufotes (Bufonidae). Izvestiya Vysshykh Uchebnykh Zavedenii. Povolzhskii Region. Natural Sciences, № 1 (25): 80–93. https://doi.org/10.21685/2307-9150-2019-1-9 (in Russian, with English summary)
- Dufresnes C., Mazepa G., Jablonski D., Oliveira Ricardo C., Wenseleers T., Shabanov D.A., Auer M., Ernst R., Koch C., Ramírez-Chaves H.E., Mulder K.P., Simonov E., Tiutenko A., Kryvokhyzha D., Wennekes P.L., Zinenko O.I., Korshunov O.V., Al-Johany A.M., Peregontsev E.A., Betto-Colliard C., Denoël M., Borkin L.J., Skorinov D.V., Pasynkova R.A., Mazanaeva L.F., Rosanov J.M., Dubey S., Litvinchuk S. 2019. Fifteen shades of green: the evolution of Bufotes toads revisited. Molecular Phylogenetics and Evolution, 141, article 106615: 1–25. doi.org/10.1016/j.ympev.2019.106615
- Borkin L.J., Andreev A.V., Litvinchuk S.N., Neupokoeva N.I., Sapelko T.V. 2019. Multidisciplinary expedition of the St. Petersburg Association of Scientists & Scholars to Nepal and Tibet (Summer 2018). Biota i Sreda Zapovednikov Dalnego Vostoka, Vladivostok, 3 (18): 124–137. https://doi.org/10.25808/26186764.2019.18.3.009 (in Russian, with English summary)
- Borkin L.J., Litvinchuk S.N., Melnikov D.A. 2020. Amphibians and reptiles of central Nepal. Pp. 123–150. In: Albedil M.F. & Borkin L.J. (eds.). Nepal: a View from Russia. St. Petersburg: Peterburgskoe Vostokovedenie (in Russian).
- Svinin A.O., Dedukh D.V., Borkin L.J., Ermakov O.A., Ivanov A.Y., Litvinchuk J.S., Zamaletdinov R.I., Mikhaylova R.I., Trubyanov A.B., Skorinov D.V., Rosanov Yu.M., Litvinchuk S.N. 2021. Genetic structure, morphological variation, and gametogenic peculiarities in water frogs (Pelophylax) from northeastern European Russia. Journal of Zoological Systematics and Evolutionary Research, 59 (3): 646–662. https://doi.org/10.1111/jzs.12447
- Borkin L.J., Andreev A.V., Vershinin V.L., Vershinina S.D., Vinarski M.V., Lopatina E.B., Neupokoeva N.I. 2021. An interdisciplinary expedition of the St. Petersburg Association of Scientists & Scholars to the Garhwal Himalaya, India (2019): some preliminary results. Biota and Environment of Natural Areas, Vladivostok, no. 1: 106–145 (in Russian, with English abstract). https://doi.org/10.37102/2782-1978_2021_1_8
- Borkin L.J., Gannibal B.K., Hochnadel V.I., Litvinchuk S.N., Moseyko A.G. 2021. Peter Simon Pallas (1741–1811) and the nature of Omsk Province. St. Petersburg: Evropeysky Dom, 111 p.
- Borkin L.J., Tikhonov A.N., Tikhonova E.P. 2021. Tibetan zoological drawings of V.I. Roborovsky, a Russian explorer of Central Asia. In: Borkin L.J. (ed.). Russian Studies on the Himalaya and Tibet - 2021: Nature and Culture. St. Petersburg: Evropeysky Dom, pp. 13–16 (in Russian).
- Borkin L.J., Baryshnikov G.F., Litvinchuk S.N., Sapelko T.V. 2021. Late Holocene mammals and reconstruction of environmental changes in the Lake Rakshastal (Langa Tso) region, Ngari Prefecture, southwestern Tibet. In: Borkin L.J. (ed.). Russian Studies on the Himalaya and Tibet - 2021: Nature and Culture. St. Petersburg: Evropeysky Dom, pp. 71–74 (in Russian).
- Borkin L.J., Litvinchuk S.N. 2021. Amphibians of the Himalaya: zoogeographical analysis. Pp. 75–78. In: Borkin L.J. (ed.). Russian Studies on the Himalaya and Tibet - 2021: the Nature and Culture. St. Petersburg: Evropeysky Dom (in Russian)
- Borkin L.J., Kovshar A.F. 2021. P.I. Rychkov (1712–1777) and the first published data on animals of Kazakhstan in the middle 18th century. Selevinia-2021, Almaty, 29: 145–165 (in Russian).
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