A current generic classification of sap beetles (Coleoptera, Nitidulidae)
A.G. Kirejtshuk
Abstract. The list of generic and subgeneric taxa arranged into subfamilies and tribes proposed for sap beetles is given, which includes also new genera [Parapocadius gen. nov. (Nitidulinae: Nitidulini) and Interfaxia gen. nov. (Nitidulinae: Cyllodini)] and subgenera [Semocarpolus subgen. nov. Gaplocarpolus subgen. nov. Askocarpolus subgen. nov. (Carpophilinae: Carpophilus Stephens, 1829)]. The Meoncerus Sharp, 1891; Apsectochilus Reitter, 1874 and Lordyrops Reitter, 1875 are considered as quite distinct each from other above mentioned as from all other generic taxa. In the list there are given the complete synonymy, including new synonymy of generic and subgeneric names [Crepuraea Kirejtshuk, 1990 and Nyujwa Perkovsky, 1990 syn. nov.; Haptoncus Murray, 1864 and Haptoncurina Jelínek, 1977, syn. nov.; Ecnomaeus Erichson, 1843 and Somaphorus Murray, 1864, syn. nov.; Ecnomorphus Motschulsky, 1858; Tribrachys LeConte, 1861 syn. nov.; Stauroglossicus Murray, 1864, syn. nov. and Microxanthus Murray, 1864, syn. nov.; Pria Stephens, 1829 and Allopria Kirejtshuk, 1980, syn. nov.; Megauchenia Macleay, 1825 and Orvoenia Dajoz, 1980, syn. nov.; Tetrisus Murray, 1864 and Pseudoischaena Grouvelle, 1897, syn. nov.; Neopocadius Grouvelle, 1906 and Pseudostelidota Grouvelle, 1906, syn. nov.; Cychramus Kugelann, 1794 and Aethinopsis Grouvelle, 1908, syn. nov.; Mystrops Erichson, 1843, and Cryptoraea Retter, 1873, syn. nov.; Cyllodes Erichson, 1843 and Mecyllodes Sharp, 1891, syn. nov.; Grammorus Murray, 1868 and Colopteroides Watrous, 1982, syn. nov.; Cryptarcha Shuckard, 1839 and Priatelus Broun, 1881, syn. nov.]. For some taxa the rank is changed, namely, Lordyra Gemminger & Harold, 1868, stat. nov. is regarded as a subgenus of Lasiodactylus Perty, 1830-1834; Brounthina Kirejtshuk, 1997, stat. nov. as a subgenus Neopocadius Grouvelle, 1906 and Teichostethus Sharp, 1891, stat. nov. as a subgenus of Hebascus Erichson, 1843, while the taxa Coxollodes Kirejtshuk, 1987 (stat. nov.) and Onicotis Murray, 1864 (stat. nov.) are regarded as separate genera. The new taxa and new taxonomical proposals are supplied with corresponding data in the notes below the list. In these notes there are also proposed the new synonymy for the following species names: Carpophilus (Ecnomorphus) acutangulus Reitter, 1884 and C. (E.) cingulatus Reitter, 1884, syn. nov.; C. (E.) bakeweli Murray, 1864; C. (E.) planatus Murray, 1864, syn. nov. and C. (E.) aterrimus Macleay, 1864, syn. nov.; C. (E.) debilis Grouvelle, 1897 and C. (E.) opaculus Grouvelle, 1897, syn. nov.; C. (E.) luridipennis Macleay, 1873 and C. (E.) loriai Grouvelle, 1906, syn. nov.; C. (E.) murrayi Grouvelle, 1892 and C. (E.) hebetatus Grouvelle, 1908, syn. nov.; C. (E.) plagiatipennis (Motschulsky, 1858) and C. (E.) nigricans Grouvelle, 1897, syn. nov.; C. (E.) terminalis Murray, 1864 and C. (E.) gentilis Murray, 1864, syn. nov.; Lasiodactylus brunneus Perty, 1830; L. centralis Cline et Carlton, 2004, syn. nov.; L. falini Cline et Carlton, 2004, syn. nov. and L. kelleri Cline et Carlton, 2004, syn. nov.); Pallodes opacus Grouvelle, 1906 and P. loriai Grouvelle, 1906, syn. nov.; Cyllodes fauveli Grouvelle, 1903 and Pallodes vagepunctus Grouvelle, 1903, syn. nov.; Pallodes jucundus Reitter, 1873 and Mecyllodes nigropictus Sharp, 1891, syn. nov.; Pallodes birmanicus Grouvelle, 1892 and P. kalingus Kirejtshuk, 1987, syn. nov.; P. gestroi Grouvelle, 1906 and P. misellus Grouvelle, 1906, syn. nov.; P. ruficollis Reitter, 1873 and P. cyanescens Grouvelle, 1898, syn. nov.; Grammophorus caelatus Gerstäcker, 1864 and Colopterus striaticollis Murray, 1864, syn. nov. In connection with a preliminary revision of many type series of the family and the mentioned taxonomical changes for some species names are established new taxonomical interpretation, namely: Pleoronia nitida (Grouvelle, 1898), comb. nov. (Axyra : Axyrodes); Parapocadius immerizi (Grouvelle, 1899), comb. nov. (Pallodes); Camptodes ruficornis (Grouvelle, 1898), comb. nov. (Pallodes); Neopallodes aestimabilis (Grouvelle, 1906), comb. nov. (Pallodes); N. alluaudi (Grouvelle, 1899), comb. nov. (Pallodes); N. aterrimus (Grouvelle, 1906), comb. nov. (Pallodes); N. dorsalis (Grouvelle, 1896), comb. nov. (Pallodes); N. fairmairei (Grouvelle, 1906), comb. nov. (Pallodes); N. incertus (Grouvelle, 1906), comb. nov. (Pallodes); N. klugi (Grouvelle, 1896), comb. nov. (Pallodes); N. limbicollis (Reitter, 1880), comb. nov. (Pallodes); N. militaris (Grouvelle, 1906), comb. nov. (Pallodes); N. niger (Grouvelle, 1906), comb. nov. (Pallodes); N. nigrocyaneus Grouvelle, 1906), comb. nov. (Pallodes); N. nitidus (Grouvelle, 1906), comb. nov. (Pallodes); N. orthogonus (Grouvelle, 1906), comb. nov. (Pallodes); N. perrieri (Grouvelle, 1906), comb. nov. (Pallodes); N. scutellaris (Grouvelle, 1906), comb. nov. (Pallodes); N. sicardi (Grouvelle, 1906), comb. nov. (Pallodes); N. sikordi (Grouvelle, 1896), comb. nov. (Pallodes); N. variabilis (Grouvelle, 1896) , comb. nov. (Pallodes); Coxollodes cyrtusoides (Reitter, 1884), comb. nov. (Pallodes); C. amamiensis (Hisamatsu, 1956), comb. nov. (Pallodes); Coxollodes opacus (Grouvelle, 1906), comb. nov. (Pallodes); C. parvulus (Grouvelle, 1908), comb. nov. (Pallodes); C. reitteri Kirejtshuk, 1987, comb. nov. (Pallodes); Pallodes fauveli (Grouvelle, 1903), comb. nov. (Cyllodes); Cyllodes jucundus (Reitter, 1873), comb. nov. (Pallodes); Interfaxia fasciata (Sharp, 1891), comb. nov.; Onicotis auritus Murray, 1864 comb. nov.; Platyarcha biguttata (Motschulsky, 1858), comb. nov. (Carphophilus: Ecnomorphus); Cryptarcha optanda (Broun, 1881), comb. nov. (Priateles). Besides, because of the new interpretation for Pallodes laetus Grouvelle, 1898, which should be transfered to the genus Camptodes, C. grouvellei nom. nov. (non Camptodes laetus Kirsch, 1873) is proposed. For the generic names Perilopa Erichson, 1843 and Meoncerus Sharp, 1891 the type species are designated as well as for Pallodes laetus Grouvelle, 1898 (Camptodes grouvellei nom. nov.) the lectotype designation is made.
Zoosystematica Rossica, 2008, 17(1): 107-122 ▪ Published in print 23 November 2008
https://doi.org/10.31610/zsr/2008.17.1.107 ▪ Open full article 
References
Cline, A.R. & Carlton, C.E. 2004. Review of Lasiodactylus Perty, with description of three new species (Coleoptera: Nitidulidae: Nitidulinae). The Coleopterists Bull., 58(3): 355-368. https://doi.org/10.1649/630
Cline, A.R. 2005. Revision of Pocadius Erichson (Coleoptera: Nitidulidae). A dissertation submitted to the Graduate Faculty of Louisiana State University and Agricultural and Mechanical College in partial fulfillment requirements for the degree of Doctor of Philosophy. http://etd.lsu.edu/docs/available/etd-11042005-122348/u, nrestricted/Cline_dis.pdf
Dajoz, R. 1980. Description de Orvoenia borneensis n. gen. n. sp., Coléoptère Colydiidae appurtenant a une tribu nouvele. Revue ent fr. (N.S,), 2(4): 190-192.
Grouvelle, A. 1908. Coléoptères de la region Indienne. Rhizodidae, Trogositidae, Nitidulidae, Colydiidae, Cucujidae. (1ere mémoire). Ann. Soc. Ent. Fr., 77: 315-495, pls.
Grouvelle, A. 1913. Byturidae, Nitidulidae. In: W. Junk (ed.). Coleopterorum Catalogus. Berlin, 15, 56: 1-223.
Hayashi, N. 1978. A contribution to the knowledge of the larvae of Nitidulidae occuring in Japan (Coleoptera: Cucujoidea). Insecta Matsum., N.S., 14: 1-97.
Jelínek, J. 1975. Redescriptions of genera Hebascus Er. And Teichostethus Sharp with designations of their type species (Coleoptera, Nitidulidae). Annatationes Zool. et Bot., 101: 1-12.
Jelínek, J. 1999. Contribution to taxonomy of the beetle subfamily Nitidulinae (Coleoptera: Nitidulidae). Folia Herovskyana, 7(5): 251-281.
Jelínek, J. & Audisio, P. 2003. Type species fixation and nomenclatural corrections in some taxa of Palaearctic Nitidulidae and Kateretidae (Coleoptera. Folia Heyrovskyana, 11(3-4): 159-171.
Jelínek, J. & Audisio, P. 2007. Nitidulidae. 459-491. In: Lobl I. & Smetana A. Catalogue of Palaearctic Coleoptera, Apollo Books, Stenstrup, 4: 935 p.
Kirejtshuk, A.G. 1986. Analysis of structure of genitalia for reconstruction of phylogeny and substantiation of system of the family of sap beetles (Nitidulidae, Coleoptera). Proc. All-Union Ent. Soc., 68: 22-28. (In Russian).
Kirejtshuk, A.G. 1987. New species of the Cyllodes complex of genera (Coleoptera, Nitidulidae) from Indochina and adjacent territoriesa. In: Entomofauna Vietnama [Entomofauna of Vietnam], Moscow, Nauka, 137-170. (In Russian).
Kirejtshuk, A.G. 1996. System, evolution of mode of life and phylogeny of the order Coleoptera. II. Ent. Obozr., 75(1): 39-62. (in Russian) [translation – 1997. System, evolution of mode of life, and phylogeny of the order Coleoptera. II. Ent. review, 76(1): 1-20].
Kirejtshuk, A.G. 1998/1999. Nitidulidae (Coleoptera) of the Himalayas and Northern Indochina. Part 1: subfamily Epuraeinae. Koeningstein, Koeltz Scientific Books (Theses Zoologicae, Vol. 28): 1-489.
Kirejtshuk, A.G. 2001. Notes on the systematics of the African Nitidulidae (Coleoptera). Ann. Hist. Nat. Mus. Nation. Hung., 93: 17-89.
Kirejtshuk, A.G. & Couturier, G. in press. Sap beetles of the tribe Mystropini (Coleoptera, Nitidulidae) associated with inflorescences of the palms cultivated in South America. Ann. Soc. Ent. Fr.
Kirejtshuk, A.G. & Jelínek, J. 2000. Preliminary review of genera of the tribe Mystropini with redescriptions and new descriptions of some genera, subgenera and species (Coleoptera: Nitidulidae: Nitidulinae). Folia Heyrovskyana, 8(3-4): 171-192.
Kirejtshuk, A.G. & Lawrence, J.F. 1992. Review of the Thalycrodes-complex of genera (Coleoptera, Nitidulidae), endemic to the Australian region. J. Austral. Ent. Soc., 31: 119-142. https://doi.org/10.1111/j.1440-6055.1992.tb00469.x
Kirejtshuk, A.G. & Lawrence, J.F. 1999. Notes on the Aethina complex (Coleoptera: Nitidulidae: Nitidulinae) with a review of the Aethina (Cleidorura) subgen.n. and Aethina (Idaethina) Gemminger & Harold. Ann. Zool., 49(3): 233-254.
Kirejtshuk, A.G. & Ponomarenko, A.G. 1990. Fossil beetles of the families Peltidae and Nitidulidae. Paleontolog. Zhurnal, 2: 78-88 + 1 pl.
Leschen, R.A.B. 1999. Systematics of Nitidulinae (Coleoptera: Nitidulidae): phylogenetic relationships, convexity and the origin of phallophagy. Invertebrate Taxonomy, 13: 845-882. https://doi.org/10.1071/IT98016
Motschulsky, V. 1858. Etudes Entomologiques. Helsingfors, 7: 192 pp. + Taf. 2.
Murray, A. 1864. Monograph of the family Nitidulidulariae. Trans. Linn. Soc. London, 24: 212-414, pls. 32-36. https://doi.org/10.1111/j.1096-3642.1863.tb00163.x
Pal, T.K. and Lawrence, J.F. 1986. A new genus and subfamily of the mycophagous Bothrideridae (Coleoptera: Cucujoidea) from the Indo-Australian region, with notes on related families. Journal of Australian Entomological Society, 25(3): 185-210. https://doi.org/10.1111/j.1440-6055.1986.tb01101.x
Parsons, C.T. 1943. A revision of Nearctic Nitidulidae (Coleoptera). Bull. Mus. Compar. Zoöl., 92(3): 121-278 + 13 pls.
Perkovsky, E.E. 1990. First discovering of Cretaceous insects of the family Leiodidae (Coleoptera) Paleontolog. Zhurnal, 4: 118-120. (In Russian).
Ponomarenko, A.G. & Kirejtshuk, A.G. 2008. Taxonomic list of fossil beetles of suborder Scarabaeina (Part 3) http://www.zin.ru/Animalia/Coleoptera/eng/paleosys2.htm (April 2008).
Sharp, D. 1891. Fam. Nitidulidae. 265-388, 8-12 tabs. In: Sharp D., Matthews A. & Lewis G. 1887-1905. Biologia Centrali-Americana. Insecta. Coleoptera, 2(1): 717 p., 19 tabs.
Spornraft, K. 1971. Zwei neue Arten der Gattung Oxycnemus Er. Und Bestimmungtabelle für die bisher bekannten Arten (Coleoptera, Nitidulidae). Opucula Zool., 116: 1-10.
