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J. Tan et al. New fossil species of ommatids (Archostemata) from the Middle Mesozoic of China...

Андрей Лобанов

J. Tan, Y. Wang, D. Ren, X. Yang
New fossil species of ommatids (Coleoptera: Archostemata) from the Middle Mesozoic of China illuminating the phylogeny of Ommatidae.
BMC Evolutionary Biology, 2012, 12:113 - doi:10.1186/1471-2148-12-113.

Authors addresses: 

Jingjing Tan ( 
Yongjie Wang ( 
Dong Ren ( 
Xingke Yang ( 

ISSN 1471-2148 

Article type Research article 
Submission date 20 February 2012 
Acceptance date 28 June 2012 

Publication date 9 July 2012 
Article URL 


Ommatidae is arguably the "most ancestral" extant beetle family. Recent species
of this group are only found in South America and Australia, but the fossil 
record reveals a much broader geographical distribution in the Mesozoic. Up to 
now, thirteen fossil genera with more than 100 species of ommatids have been 
described. However, the systematic relationships of the extant and extinct 
Ommatidae have remained obscure. Three constraint topologies were designed 
based on Kirejtshuk's hypothesis, enforced the monophyly of 
Tetraphalerus + Odontomma, Pareuryomma + Notocupes and both respectively. 

In this study, four new species, Pareuryomma ancistrodonta sp. nov., 
Pareuryomma cardiobasis sp. nov., Omma delicata sp. nov., and Tetraphalerus 
decorosus sp. nov., are described. Based on well-preserved fossil specimens and
previously published data the phylogenetic relationships of extant and extinct 
lineages of Ommatidae were analyzed for the first time cladistically. Based on 
the results we propose a new classification with six tribes of Ommatidae: 
Pronotocupedini, Notocupedini, Lithocupedini, Brochocoleini, Ommatini and 
Tetraphalerini. These taxa replace the traditional four subfamilies. 

There is good support for the monophyly of the ingroup. Notocupedini, as 
defined by Ponomarenko, are paraphyletic. Notocupoides + Eurydictyon are the 
sister group of the remaining fossil and extant ommatids. Together they form 
the clade Pronotocupedini. 
Notocupedini and Lithocupedini are the next two branches. The tribe 
Brochocoleini is the sister group of a clade comprising Tetraphalerini and 


Ommatidae, Omma, Tetraphalerus, New species, Mesozoic, China 

Файл PDF: tan_et_al_ommatids_2012.pdf