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И.В. Шохин. 2019.
Фауна пластинчатоусых жуков (Coleoptera: Scarabaeoidea) Азербайджана.
Кавказский энтомологический бюллетень, 2019, 15(1): 61-106.
[I.V. Shokhin. The fauna of lamellicorn beetles (Coleoptera: Scarabaeoidea)
of Azerbaijan / Caucasian Entomological Bulletin, 2019, 15(1): 61-106.]
DOI: 10.23885/181433262019151-61106
Резюме.
На основании литературных и
коллекционных данных впервые составлен список
надсемейства Scarabaeoidea фауны Азербайджана.
Список почти в 2 раза увеличивает количество
отмеченных в страны таксонов: если в 2004 году
их число оценивалось в 162 вида, то к настоящему
времени зарегистрировано не менее 298 видов и
12 подвидов, кроме того, еще 4 вида известны с этой
территории только по первоописанию; 33 таксона
обитают в пограничных районах и могут быть
найдены в Азербайджане. Статус 16 таксонов требует
пересмотра. Разработана оригинальная концепция
полиморфного вида Protaetia (Netocia) hieroglyphica
(Menetries, 1832), sensu nova представленного на
территории Азербайджана 4 подвидами: Protaetia
(Netocia) hieroglyphica hieroglyphica (Menetries, 1832),
P. h. cuprina (Motschulsky, 1849), sensu nova, stat. n.,
P. h. alboflagellata (Olsoufieff, 1916), subsp. resurr.,
sensu nova, P. h. caucasica (Kolenati, 1846), sensu nova,
stat. n. Азербайджанская популяция Glaphyrus
oxypterus Pallas, 1771 выделена в отдельный подвид
Glaphyrus oxypterus kasatkini subsp. n. Предложена
новая синонимия: Decamera Mulsant, 1842 =
Odonthoplia Zaitsev, 1924, syn. n.; Melolontha persica
Reitter, 1902, nom. resurr. = Melolontha kraatzi Reitter,
1906 (замещающее название для Melolontha tibialis
Kraatz, 1882, non Mulsant, 1842), syn. n. = Melolontha
tibialis var. brenskei Reitter, 1902, syn. n.; Trypocopris
fausti (Reitter, 1890) = Geotrupes zaitzevi Olsoufieff,
1918, syn. n., Protaetia (Netocia) araratica (Reitter,
1891) = Potosia schelkovnikovi Zaitzev, 1918, syn. n.
Предложено понизить статус рода Glyptotrox Nikolajev,
2016, stat. n. до подрода в составе рода Trox Fabricius,
1775. Последние указания для Азербайджана таких
видов, как Onthophagus kolenatii Reitter, 1892 (синоним
Onthophagus (Palaeonthophagus) nuchicornis (Linnaeus,
1758)) и Onthophagus basipustulatus Heyden, 1889
(данные относятся к Onthophagus (Palaeonthophagus)
formaneki Reitter, 1897) являются ошибочными.
В список включен ряд видов, известных из пограничных
с Азербайджаном районов.
Abstract.
A checklist of the Scarabaeoidea of
Azerbaijan is given for the first time. In total 298 species
and 12 subspecies of scrarab beetles are registered
for Azerbaijan, from which Sinodendron cylindricum
(Linnaeus, 1758), Bodilus insperatus (Petrovitz, 1967) and
Amphimallon circassicum Brenske, 1894 are recorded for
the country for the first time. In addition 4 species are
known only by the original descriptions. The following
species are distributed in border areas of Russia, Georgia,
Armenia, Iran, and can be found in Azerbaijan in future
or these taxa have unclear status: Dorcus peyronis
Reiche, Saulcy, 1856, Trox (Granulitrox) hispidus niger
Rossi, 1792, Brenskea coronata Reitter, 1891, Eulasia
(s. str.) praeusta (Champenois, 1896), Eulasia (s. str.)
azarbaijanica (Petrovitz, 1980), Eulasia (s. str.) aurantica
(Reitter, 1890), Eulasia (s. str.) corniculata (Reitter, 1903),
Pygorpleurus gordyenensis (Petrovitz, 1971), Glaphyrus
varians Menetries, 1836, Scarabaeus sacer Linnaeus, 1758,
Scarabaeus babori Balthasar, 1934б Scarabaeus wilsoni
C.O. Waterhouse, 1890, Onthophagus (Palaeonthophagus)
excubitor Ziani et Gudenzi, 2006, Onthophagus
(Palaeonthophagus) flagrans Reitter, 1892, Pararhyssemus
coluber Mayet, 1887, Bodilus circumcinctus (W. Schmidt,
1840), Neagolius abchasicus (Reitter, 1892), Biralus equinus
(Faldermann, 1835), Chilothorax conspurcatus (Linnaeus,
1758), Chilothorax pictus (Sturm, 1805), Chilothorax
lineolatus (Illiger, 1803), Melinopterus consputus
(Creutzer, 1799), Sigorus porcus (Fabricius, 1792), Pubinus
tomentosus (Muller, 1776), Esymus suturinigra (Schmidt,
1916), Euorodalus paracoenosus (Balthasar et Hrubant,
1960), Phalacronotus quadrimaculatus (Linnaeus,
1761), Mendidius bispinifrons (Reitter, 1889), Mendidius
multiplex (Reitter, 1897), Planolinus borealis Gyllenhal,
1827, Planolinus fasciatus Olivier, 1789, Agoliinus satyrus
(Gyllenhal, 1808), Agoliinus piceus (Gyllenhal, 1808),
Pseudoesymus lucidus (Klug, 1845), Polyphylla (s. str.)
fullo (Linnaeus, 1758), Cryptotrogus maluzhenkoi (Zaitzev,
1928), Holochelus (Eriotrogus) erivanicus (Reitter, 1902),
Holochelus (Eriotrogus) majusculus (Nonveiller, 1965),
Amphimallon altaicum (Mannerheim, 1825), Amphimallon
arnoldii (Medvedev, 1951), Lasiopsis canina (Zoubkov,
1829), Hoplia (s. str.) hyrcana Medvedev, 1952, Hoplia
(s. str.) parvula Krynicki, 1832, Phyllopertha horticola
(Linnaeus, 1758), Anisoplia (s. str.) agricola (Poda, 1761),
Anisoplia (s. str.) parva Kraatz, 1883, Pentodon foveipennis
Jakovlev, 1903, Pentodon bidens bidens (Pallas, 1771),
Osmoderma richteri Medvedev, 1953.
The original concept of polymorphic Protaetia
(Netocia) hieroglyphica (Menetries, 1832), sensu nova
species-complex, presented in the territory of Azerbaijan
by 4 subspecies, was proposed. Here we distinguish the
hieroglyphica-complex, comprising taxa with white spots
on the elytra and prosternum, in contrast to the cuprea-
complex with mainly Western monochrome (without white
spots) species (cuprea Fabricius, 1775, obscura Andersch,
1797, olivacea Mulsant, 1842). This new complex includes
four close subspecies (possibly forming hybrids), that differ
in morphology, distribution and bionomics: Protaetia
(Netocia) hieroglyphica hieroglyphica (Menetries, 1832),
P. h. cuprina (Motschulsky, 1849), sensu nova, stat. n.,
P. h. alboflagellata (Olsoufieff, 1916), subsp. resurr., sensu
nova, P. h. caucasica (Kolenati, 1846), sensu nova, stat. n.
A detailed revision of all Caucasian taxa of cuprina species-
group will be published in a separate paper.
A new subspecies is described for Azerbaijanian
population of Glaphyrus oxypterus Pallas, 1771: Glaphyrus
oxypterus kasatkini subsp. n. The new subspecies differs
from the nominotypical one by the rounded apex of the
male elytra (angle-shaped in the nominotypical subspecies).
This character is similar to those in Glaphyrus caucasicus
Kraatz, 1882. The following differences are found in the
structure of the endophallus: the dorsal camera is with
constriction and widened to apex in the new subspecies,
but without constriction and with straight sides in the
nominotypical subspecies; the agonoporium on the frontal
camera forms small tooth-like protrusion in the new
subspecies, but it is located on an elongate and bent down
process in the nominotypical subspecies.
The following new synonymy is proposed: Melolontha
persica Reitter, 1902, nom. resurr. = Melolontha kraatzi
Reitter, 1906 (replacement name for Melolontha tibialis
Kraatz, 1882 non Mulsant, 1842), syn. n. = Melolontha
tibialis var. brenskei Reitter, 1902, syn. n.; Trypocopris
fausti (Reitter, 1890) = Geotrupes zaitzevi Olsoufieff,
1918, syn. n.; Protaetia (Netocia) araratica (Reitter,
1891) = Potosia schelkovnikovi Zaitzev, 1918, syn. n. The
status of the genus Glyptotrox Nikolajev, 2016, stat. n. is
changing to a subgenus in the genus Trox Fabricius, 1775.
The following two species are erroneously recorded for the
fauna of Azerbaijan: Onthophagus kolenatii Reitter, 1892
(the junior synonym of Onthophagus (Palaeonthophagus)
nuchicornis (Linnaeus, 1758)) and Onthophagus
basipustulatus Heyden, 1889 (data refer to Onthophagus
(Palaeonthophagus) formaneki Reitter, 1897).