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A. Bezděk, J. Hájek. 2012.
Catalogue of type specimens of beetles (Coleoptera) deposited in the National Museum, Prague, Czech Republic. Scarabaeoidea (part V). Scarabaeidae: Scarabaeinae: Coprini, Eurysternini, Gymnopleurini and Oniticellini.
Acta Entomologica Musei Nationalis Pragae, 2012, 52 (1): 297-334.