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C.J. Wilson & R.D. Angus. 2004.
A chromosomal analysis of ten European species of Aphodius Illiger,
subgenera Acrossus Mulsant, Nimbus Mulsant & Rey and Chilothorax Motschulsky (Coleoptera: Aphodiidae).
Koleopterologische Rundschau, 2004, 74: 367-374.
This paper (C. Wilson and R. Angus) is avaliable here: PDF (244 kb).
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