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Classification of extinct and recent Coleoptera

Curator of project - A.G. Kirejtshuk

(besides the author and year for most taxa
the codes of geochronologic divisions and marks
on distribution in the Palaearctic region and Russia are included)


 Ordo: COLEOPTERA Linnaeus [ ... ] Pal/Rus
 =  Scarabaeida Laicharting, 1781
+Subordo: Protocoleoptera [ ... ]
 =  Archecoleoptera
+Superfamilia: Tshekardocoleoidea Rohdendorf, 1944 [ ... ] Pal/Rus
+Familia: Tshekardocoleidae Rohdendorf, 1944 [P1ar ... P2kz] Pal/Rus
+Familia: Labradorocoleidae Ponomarenko, 1969 [Kal3-cm1 ... Kal3-cm1]
+Superfamilia: Permocupedoidea Martynov, 1933 [P2uf ... T3k] Pal/Rus
+Familia: Permocupedidae Martynov, 1933 [P2uf ... P2t] Pal/Rus
 =  Kaltanocoleidae Rohdendorf, 1961
+Familia: Taldycupedidae Rohdendorf, 1961 [P2kz ... T3k] Pal/Rus
 Subordo: Archostemata Kolbe, 1908 [ ... ] Pal/Rus
 =  Cupedina
 Infraordo: Cupediformia {Kirejtshuk, 1991} [ ... ] Pal/Rus
 =  Cupedomorpha ?
 Superfamilia: Cupedoidea Laporte, 1836 [ ... ] Pal/Rus
 Familia: Cupedidae Laporte, 1836 [T21 ... R] Pal/Rus
 =  Brochocoleidae Hong, 1982
 =  Cupesidae Lacordaire, 1858
 Familia: Crowsoniellidae Iablokoff-Khnzorian, 1983 [ ... R] Pal/-
 Familia: Ommatidae Sharp et Muir, 1912 [J1he ... R] Pal/Rus
 =  Ommadidae
 =  Tetraphaleridae
 Subfamilia: Ommatinae Sharp et Muir, 1912 [??? ... R] Pal/Rus
 Subfamilia: Tetraphalerinae Crowson, 1962 [??? ... R] Pal/Rus
+Familia: Triadocupedidae Ponomarenko, 1966 [T2-3l-c ... ???] Pal/?
 Familia: Jurodidae Ponomarenko, 1985 [J12 ... R] Pal/Rus
 =  Sikhotealiniidae Lafer, 1996
 Infraordo: Micromaltiformia Kirejtshuk, 1991 [ ... R] Pal/-
 Superfamilia: Micromalthoidea ?????? [K1ap, Leba ... R] Pal/-
 Familia: Micromalthidae Barber, 1913 [K1ap, Leba ... R] Pal/-
 Subordo: INCERTAE SEDIS: [ ... ]
+Superfamilia: Ademosynoidea Ponomarenko, 1969 [ ... ]
+Familia: Ademosynidae Ponomarenko, 1969 [P2t ... K12] Pal/?
 Subordo: Polyphaga Emery, 1886 [ ... ] Pal/Rus
 =  Scarabaeida
 Infraordo: Staphyliniformia Lameere, 1900 [ ... ] Pal/Rus
 =  Scarabaeiformia ???
 =  Scarabaeomorpha
 =  Staphylinomorpha
 Superfamilia: Hydrophyloidea Latreille, 1802 [ ... ] Pal/Rus
 Familia: Hydraenidae Mulsant, 1844 [J11 ... R] Pal/Rus
 =  Limnebiidae Mulsant, 1844
 =  Ochtebiidae C.G. Thomson, 1859
 Subfamilia: Prostethopinae Perkins in Perkins et Balfour-Browne, 1994 [ ... R] ?/?
 Subfamilia: Hydraeninae Mulsant, 1844 [ ... R] Pal/Rus
 Subfamilia: Ochthebiinae C.G. Thomson, 1859 [Pg1d ... R] Pal/Rus
 Familia: Hydrochidae C.G. Thomson, 1859 [Pg22 ... R] Pal/Rus
 Familia: Helophoridae Leach, 1815 [N13 ... R] Pal/Rus
 =  Elophorinae
 Familia: Spercheidae Erichson, 1837 [ ... R] Pal/Rus
 Familia: Hydrophilidae Latreille, 1802 [J11 ... R] Pal/Rus
 =  Sphaeridiidae Latreille, 1802
 Subfamilia: Epimetopinae Zaitzev, 1908 [ ... R] ?/?
 Subfamilia: Horelophinae Hansen, 1991 [ ... R] -/-
 Subfamilia: Sphaeridiinae Latreille, 1802 [ ... R] Pal/Rus
 Subfamilia: Hydrobiinae Mulsant, 1844 [ ... R] Pal/Rus
 Subfamilia: Chetarthriinae [ ... R] ?/?
 Subfamilia: Horelophinae Hansen, 1991 [ ... R] Pal/Rus
 Subfamilia: Hydrophilinae Latreille, 1802 [ ... R] Pal/Rus
 =  Amphiopinae Orchym., 1919
 Subfamilia: Berosinae Mulsant, 1844 [Pg22, GreR ... R] Pal/Rus
 Familia: Georissidae Laporte, 1819 [Pg22 ... R] Pal/Rus
 =  Georyssidae Laporte, 1819
 Superfamilia: Histeroidea Gyllenhal, 1808 [ ... ] Pal/Rus
 Familia: Sphaeritidae Shuckard, 1839 [ ... R] Pal/Rus
 Familia: Synteliidae Lewis, 1882 [ ... R] Pal/Rus
 Familia: Histeridae Gyllenhal, 1808 [Pg22 ... R] Pal/Rus
 =  Niponiidae Handlirsch, 1925
 Subfamilia: Dendrophilinae Reitter, 1909 [Pg22, Mess ... R] Pal/Rus
 Subfamilia: Hetaeriinae Marseul, 1857 [ ... R] Pal/Rus
 Subfamilia: Histerinae Gyllenhal, 1808 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Onthophilinae MacLeay, 1819 [ ... R] Pal/Rus
 Subfamilia: Tribalinae Bickhardt, 1914 [ ... R] Pal/Rus
 Subfamilia: Abraeinae MacLeay, 1819 [ ... R] Pal/Rus
 Subfamilia: Chlamydopsinae Bickhardt, 1914 [ ... R] ?/?
 Subfamilia: Niponiinae Fowler, 1912 [ ... R] ?/?
 Subfamilia: Saprininae Blanchard, 1845 [ ... R] Pal/Rus
 Subfamilia: Trypanaeinae Marseul, 1857 [ ... R] ?/?
 =  Triponaeinae
 Subfamilia: Trypeticinae Bickhardt, 1914 [ ... R] ?/?
 Superfamilia: Staphylinoidea Latreille, 1802 [ ... ] Pal/Rus
 Familia: Agyrtidae C.G. Thomson, 1859 [J12 ... R] Pal/Rus
 =  Silphidae, part.
 Subfamilia: Lyrosominae Horn, 1880 [ ... R] ?/?
 Subfamilia: Pterolominae Port., 1914 [ ... R] Pal/Rus
 Subfamilia: Catopocerinae Hatch, 1927 [ ... R] ?/?
 =  Glacicavicolinae
 =  Pinodytinae Horn, 1880
 Familia: Colonidae Horn, 1880 [ ... R] Pal/Rus
 =  Leptinidae LeConte, 1866
 =  Platypsilidae Ritsema, 1869
 Subfamilia: Coloninae Horn, 1880 [ ... ] Pal/Rus
 Subfamilia: Platypsilinae Ritsema, 1869 [ ... R] Pal/Rus
 Subfamilia: Leptininae LeConte, 1866 [ ... R] Pal/Rus
 Familia: Leiodidae Fleming, 1821 [K12 ... R] Pal/Rus
 =  Agathidiidae Westwood, 1839
 =  Anisotomidae Erichson, 1845
 =  Camiaridae Jeannel, 1911
 =  Catopidae Jeannel, 1922
 =  Cholevidae Jeannel, 1922
 =  Leptodiridae Abeille de Perrin, 1878
 =  Liodidae Jacobson, 1910
 =  Sogdiidae
 Subfamilia: Camiarinae Jeannel, 1911 [ ... R] ?/?
 =  Agyrtodinae
 Subfamilia: Liodinae Fleming, 1951 [ ... R] Pal/Rus
 =  Anisotominae
 Subfamilia: Cholevinae Kirby, 1837 [K12 ... R] Pal/Rus
 =  Anemadinae Jeannel, 1922
 =  Bathysciinae Horn, 1880
 =  Catopinae Jeannel, 1922
 =  Eucatopinae Jeannel, 1921
 =  Leptodirinae Abeille de Perrin, 1878
 =  Nemadinae
 =  Ptomaphaginae
 Familia: Silphidae Latreille, 1807 [Pg23 ... R] Pal/Rus
 Subfamilia: Nicrophorinae Kirby, 1837 [Pg2-3 ... R] Pal/Rus
 Subfamilia: Silphinae Latreille, 1802 [ ... R] Pal/Rus
 Familia: Scydmaenidae Leach, 1815 [K12 ... R] Pal/Rus
 =  Anisosphaeridae Temoesvary, 1882
 Subfamilia: Mastiginae Fleming, 1821 [ ... R] ?/?
 Subfamilia: Scydmaeninae Leach, 1815 [ ... R] Pal/Rus
 Familia: Ptiliidae Heer, 1843 [Pg23 ... R] Pal/Rus
 =  Cephaloplectidae Sharp, 1882
 =  Limulodidae Ganglbauer, 1899
 =  Trichopterigidae Erichson, 1845
 Subfamilia: Acrotrichinae Reitter, 1909 [ ... R] Pal/Rus
 Subfamilia: Cephaloplectinae Sharp, 1883 [ ... R] ?/?
 =  Limulodinae Ganglbauer, 1899
 Subfamilia: Nanosellinae Barber, 1924 [ ... R] ?/?
 Subfamilia: Ptiliinae Erichson, 1845 [ ... R] Pal/Rus
 Familia: Staphylinidae Latreille, 1802 [J12 ... R] Pal/Rus
 =  Brathinidae
 =  Dasyceridae Reitter, 1887
 =  Empelidae
 =  Micropeplidae Leach, 1815 ?(Heer, 1838)
 =  Oxyporidae Fleming, 1821
 =  Oxytelidae Fleming, 1821
 Subfamilia: Glypholomatinae Jeannel, 1962 [ ... ] -/-
 Subfamilia: Microsilphinae Crowson, 1950 [ ... R] -/-
 Subfamilia: Dasycerinae Reitter, 1887 [ ... R] Pal/Rus
 Subfamilia: Protopselaphinae Newton et Thayer, 1995 [ ... R] -/-
 Subfamilia: Pselaphinae Latreille, 1802 [Pg23 ... R] Pal/Rus
 =  Batrisinae Reitter, 1882 ?(Raffrai, 1890)
 =  Bythininae Raffrai, 1890
 =  Clavigerinae Leach, 1815
 =  Euplectinae
 =  Faroninae Reitter, 1882
 =  Goniacerinae Reitter, 1882
 =  Mayetiinae
 =  Pselaphinae Latreille, 1802
 Subfamilia: Trichophyinae C.G. Thomson, 1858 [ ... R] -/-
 Subfamilia: Phloeocharinae Erichson, 1839 [ ... R] Pal/Rus
 Subfamilia: Scaphidiinae Latreille, 1806 [Pg23 ... R] Pal/Rus
 Subfamilia: Piestinae Erichson, 1839 [ ... R] -/-
 Subfamilia: Aphaenostemminae Peyerimhoff, 1914 [ ... R] ?/?
 Subfamilia: Empelinae (Abdullah, 1969) Newton et Thayer, 1991 [ ... R] -/-
 Subfamilia: Omaliinae Macleay, 1825 [J13 ... R] Pal/Rus
 =  Brathinidae
 =  Aphaenostemminae
 Subfamilia: Proteininae Erichson, 1839 [ ... R] Pal/Rus
 =  Metopsiinae
 =  Pteroniinae
 Subfamilia: Micropeplinae Leach, 1815 [ ... R] Pal/Rus
 Subfamilia: Trilobitideinae Eichelbaum, 1909 [ ... ]
 Subfamilia: Pseudopsinae Ganglbauer, 1895 [ ... R] ?/?
 Subfamilia: Olisthaerinae C.G. Thomson, 1859 [J21 ... R] Pal/Rus
 Subfamilia: Solieriinae Newton et Thayer, 1991 [ ... R] -/-
 Subfamilia: Apateticinae Fauvel, 1895 [ ... R] ?/?
 Subfamilia: Trigonurinae Reiche, 1845 [J33 ... R] Pal/Rus
 Subfamilia: Osoriinae Erichson, 1839 [Pg23, BalJ ... R] ?/?
 Subfamilia: Neophoninae Fauvel, 1905 [ ... R] ?/?
 Subfamilia: Habrocerinae Mulsant et Rey, 1877 [ ... R] Pal/Rus
 Subfamilia: Aleocharinae Fleming, 1821 [ ... R] Pal/Rus
 =  Cyphinae
 =  Hypocyptinae Ganglbauer, 1895
 =  Mesoporinae
 =  Mimanommatinae Wasmann, 1912
 =  Pseudoperinthinae
 =  Pulicomorphinae Mann, 1921
 =  Pygosteninae Wasmann, 1902
 =  Termitodiscinae Wasmann, 1912
 =  Trichopseniinae Eichelbaum, 1909
 Subfamilia: Oxyporinae Fleming, 1821 [ ... R] Pal/Rus
 Subfamilia: Oxytelinae Fleming, 1821 [ ... R] Pal/Rus
 Subfamilia: Megalopsidiinae Leng, 1920 [ ... R] -/-
 Subfamilia: Steninae Macleay, 1825 [ ... R] Pal/Rus
 Subfamilia: Euaesthetinae C.G. Thomson, 1859 [ ... R] ?/?
 Subfamilia: Leptothyphlinae Fauvel, 1874 [ ... R] ?/?
 Subfamilia: Trichophyinae C.G. Thomson, 1859 [ ... R] -/-
 Subfamilia: Tachyporinae MacLeay, 1825 [J31, KarT ... R] Pal/Rus
 Subfamilia: Paederinae Ganglbauer, 1895 [Pg22, GreR ... R] Pal/Rus
 Subfamilia: Staphylininae Latreille, 1802 [ ... R] Pal/Rus
 =  Amblyopininae
 =  Diochinae
 =  Platyprosopinae
 =  Quedinae Ganglbauer, 1895
 =  Xantholininae Ganglbauer, 1895
 =  Xanthoperiginae
 Superfamilia: Scarabaeoidea Latreille, 1802 [ ... ] Pal/Rus
 =  Lamellicornia
 Familia: Glaresidae Semenov-Tian-Shanskii et Medvedev, 1932 [ ... R] Pal/?
 Familia: Passalidae Leach, 1815 [ ... R] -/-
 Subfamilia: Aulacocyclinae Kaup, 1868 [Pg33 ... R] -/-
 Subfamilia: Passalinae Leach, 1815 [Post, N12, Sha ... R] -/-
 Familia: Lucanidae Latreille, 1804 [K2t ... R] Pal/Rus
 Subfamilia: Aesalinae Latreille, 1804 [ ... R] Pal/Rus
 Subfamilia: Nicaginae LeConte, 1861 [ ... R] Pal/?
 Subfamilia: Syndesinae MacLeay, 1819 [Pg31 ... R] Pal/Rus
 =  Sinodendrinae
 Subfamilia: Lampriminae MacLeay, 1819 [Pg23, BalJ ... R] -/-
 Subfamilia: Penichrolucaninae Arrow, 1950 [ ... R] -/-
 Subfamilia: Lucaninae Latreille, 1804 [Pg23 ... R] Pal/Rus
 =  Chiasognathinae
 =  Dorcinae
 =  Figulinae
 =  Odontolabinae
 =  Platycerinae
 Familia: Trogidae Macleay, 1819 [ ... ] Pal/Rus
 Familia: Pleocomidae LeConte, 1861 [ ... R] -/-
 Familia: Diphyllostomatidae Holloway, 1972 [ ... R] -/-
 Familia: Geotrupidae Latreille, 1802 [K12 ... R] Pal/Rus
 Subfamilia: Bolboceratinae Mulsant, 1842 [K12 ... R] ?/?
 =  Athyreinae Howden et Martinetz, 1963
 =  Bolbocerinae
 Subfamilia: Geotrupinae Latreille, 1802 [ ... R] Pal/Rus
 =  Taurocerastinae
 Subfamilia: Lethrinae Mulsdant et Rey, 1871 [ ... R] Pal/Rus
 Familia: Ochodaeidae Mulsant et Rey, 1871 [K1ap ... R] Pal/Rus
 Subfamilia: Ochodaeinae Mulsant et Rey, 1871 [ ... R] Pal/Rus
 Subfamilia: Chaetocanthinae Scholtz in Scholtz et al., 1988 [ ... R] -/-
 Familia: Belohinidae Paulian, 1959 [ ... R] ?/?
 Familia: Ceratocanthidae Cartwright et Gordon, 1971 [ ... R] -/-
 =  Acanthoceridae
 Familia: Glaphyridae Macleay, 1819 [ ... R] ?/?
 =  Lichniinae
 Familia: Scarabaeidae Latreille, 1802 [J2 ... R] Pal/Rus
 =  Aclopidae
 =  Cetoniidae
 Subfamilia: Lithoscarabaeinae Nikolajev, 1992 [K12, Bais ... R] Pal/?
 Subfamilia: Hybosorinae Erichson, 1847 [K12 ... R] ?/?
 Subfamilia: Aclopinae Milne-Edwards, 1850 [J3ox ... R] -/-
 Subfamilia: Scarabaeinae Latreille, 1802 [Pg1d ... R] Pal/Rus
 Subfamilia: Coprinae [K2tu ... R] Pal/Rus
 Subfamilia: Aphodiinae Leach, 1815 [K12 ... R] Pal/Rus
 =  Aegaliinae
 =  Aulonocneminae
 =  Chironinae
 =  Silluviinae
 =  Termithtroginae
 Subfamilia: Pachypodinae Erichson, 1840 [ ... R] Pal/Rus
 Subfamilia: Oncerinae [ ... R] ?/?
 Subfamilia: Orphninae Erichson, 1847 [ ... R] ?/?
 Subfamilia: Allidostomatinae Arrow, 1940 [ ... R] -/-
 =  Allidiostominae
 =  Idiostominae
 Subfamilia: Dynamopodinae Arrow, 1911 [ ... R] Pal/?
 =  Dynamopinae
 Subfamilia: Melolonthinae MacLeay, 1819 [Pg21, Wilc ... R] Pal/Rus
 =  Syntellopodinae
 Subfamilia: Sericinae [K12 ... R] Pal/Rus
 Subfamilia: Hopliinae [Pg31, Flor ... R] Pal/Rus
 Subfamilia: Euchirinae Hope, 1840 [N12, Tott ... R] Pal/?
 Subfamilia: Phenomeridinae Erichson, 1847 [ ... R] ?/?
 =  Phenomerinae
 Subfamilia: Cetoniinae Leach, 1815 [N12 ... R] Pal/Rus
 =  Trichinae
 =  Valginae
 Subfamilia: Dynastinae MacLeay, 1819 [Pg31 ... R] ?/?
 Subfamilia: Rutelinae MacLeay, 1819 [Pg22, Mess ... R] Pal/Rus
 Infraordo: Elateriformia [ ... ] Pal/Rus
 =  Dascilliformia
 =  Eucinetiformia
 Superfamilia: Dascilloidea Gue'rin-Me'neville, 1843 [ ... ] Pal/Rus
 Familia: Dascilidae Gue'rin-Me'neville, 1843 [K2cm ... R] Pal/Rus
 =  Karumiidae Escalera, 1913
 Subfamilia: Dascillinae Gue'rin-Me'neville, 1843 [K2cm ... R] Pal/Rus
 =  Atopinae
 =  Cinnabariinae
 Subfamilia: Karumiinae Escalera, 1913 [ ... R] Pal/?
 =  Zarudniolidae
 Familia: Rhipiceridae Latreille, 1834 [ ... R] Pal/Rus
 =  Sandalidae
 Superfamilia: Scirtoidea Fleming, 1821 [ ... ] Pal/Rus
 =  Clamboidea Fischer, 1821
 =  Eucinetoidea Lacordaire, 1857
 Familia: Decliniidae Nikit., Lawr., Kirejt. et Gratsh., 1994 [ ... R] Pal/Rus
 Familia: Eucinetidae Lacordaire, 1857 [J3ox ... R] Pal/Rus
 =  Cryptomeridae
 Familia: Clambidae Fischer, 1821 [Pg23 ... R] Pal/Rus
 =  Calyptomeridae
 Subfamilia: Calyptomerinae Crowson, 1955 [ ... R] ?/?
 Subfamilia: Acalyptomerinae [ ... R] ?/?
 Subfamilia: Clambinae Fischer, 1821 [ ... R] Pal/Rus
 Familia: Scirtidae Fleming, 1821 [Pg23 ... R] Pal/Rus
 =  Cyphonidae Stephens, 1829
 =  Elodidae Shuckard, 1840
 =  Helodidae Agassiz, 1846
 Superfamilia: Buprestoidea [ ... ] Pal/Rus
 Familia: Buprestidae Leach, 1815 [J2bt ... R] Pal/Rus
 =  Electrapatidae Iablokoff-Khnzorian, 1962
 Subfamilia: Schizopodinae Leconte, 1861 [ ... P] -/-
 Subfamilia: Julodinae Lacordaire, 1857 [ ... P] Pal/?
 Subfamilia: Parathyreinae Alexeev, 1993 [J2bt ... K1]
 Subfamilia: Chalcophorinae Lacordaire, 1857 [Pg22 ... R] Pal/Rus
 Subfamilia: Buprestinae Leach, 1815 [Pg13, Mena ... R] Pal/Rus
 =  Chrysobothrinae
 Subfamilia: Mastogeniinae Leconte et Horn, 1883 [Pg23 ... R]
 Subfamilia: Sphenopterinae Lacordaire, 1857 [Pg22, Mess ... R] Pal/Rus
 Subfamilia: Agrilinae Laporte, 1835 [Pg ... R] Pal/Rus
 =  Galbellinae
 Subfamilia: Polycestinae Lacordaire, 1857 [ ... P] ?/?
 Subfamilia: Acmaeoderinae [Pg13, Mena ... R] Pal/Rus
 Subfamilia: Sternocerinae [ ... ]
 Subfamilia: Thrincopyginae LeConte, 1861 [ ... R] ?/?
 Subfamilia: Cylindromorphinae Portevin, 1931 [ ... R] Pal/-
 Subfamilia: Trachyinae Laporte, 1835 [ ... R] Pal/Rus
 Superfamilia: Byrrhoidea [ ... ] Pal/Rus
 =  Dryopoidea
 =  Psephenoidea
 Familia: Byrrhidae Latreille, 1806 [J11 ... R] Pal/Rus
 =  Syncaliptidae Mulsant et Rey, 1869
 Subfamilia: Byrrinae Latreille, 1806 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Amphicyrtinae Leconte, 1861 [Pg31, Flor ... R] ?/?
 Subfamilia: Syncaliptinae Mulsant et Rey, 1869 [Pg23, BalJ ... R] ?/?
 Familia: Elmidae Curtis, 1830 [Pg22 ... R] Pal/Rus
 =  Elmididae
 =  Elminthidae
 =  Helminthidae
 =  Linmiidae
 Subfamilia: Larainae LeConte, 1861 [Pg22, Geis ... R] ?/?
 =  Larinae
 Subfamilia: Elminae Curtis, 1830 [Pg23 ... R] Pal/Rus
 Familia: Dryopidae Billberg, 1820 [Pg31 ... R] Pal/Rus
 =  Chiloeidae
 =  Parnidae
 Familia: Lutrochidae Kasap et Crowson, 1975 [Pg31 ... R] -/-
 Familia: Limnichidae Erichson, 1846 [Pg23 ... R] Pal/Rus
 Subfamilia: Hyphalinae Britton, 1971 [ ... ]
 Subfamilia: Limnichinae Erichson, 1846 [Pg23 ... R] Pal/Rus
 Subfamilia: Cephalobyrrhinae Champion, 1925 [ ... R] ?/?
 Subfamilia: Thaumastodinae Champion, 1924 [ ... R] ?/?
 Familia: Heteroceridae MacLeay, 1825 [K11 ... R] Pal/Rus
 Subfamilia: Elythomerinae Pacheco, 1964 [ ... R] ?/?
 Subfamilia: Heterocerinae MacLeay, 1825 [K11 ... R] Pal/Rus
 Familia: Psephenidae Lacordaire, 1854 [Pg22 ... R] Pal/Rus
 =  Eubriidae
 Familia: Psephenoididae [ ... ]
 Subfamilia: Eubriinae Lacordaire, 1857 [ ... R] Pal/Rus
 Subfamilia: Psephenoidinae Hinton, 1939 [ ... R] ?/?
 Subfamilia: Eubrianacinae Jacobson, 1913 [ ... R] ?/?
 =  Placonychinae Horn, 1880
 Subfamilia: Psepheninae Lacordaire, 1854 [Pg31, Flor ... R] Pal/Rus
 Familia: Cneoglossidae Champion, 1897 [ ... R] -/-
 Familia: Ptylodactylidae Laporte, 1836 [Pg23 ... R] Pal/Rus
 Subfamilia: Anchytarsinae Champion, 1897 [ ... R] Pal/-
 Subfamilia: Cladotominae Pic, 1914 [ ... R] Pal/-
 Subfamilia: Aploglossinae Champion, 1897 [ ... R] -/-
 Subfamilia: Araeopidiinae Lawrence, 1991 [ ... R] -/-
 Subfamilia: Ptylodactylinae Laporte, 1836 [Pg23 ... R] Pal/Rus
 Familia: Chelonariidae Blanchard, 1845 [Pg31 ... R] -/-
 Familia: Eulichadidae Crowson, 1973 [ ... R] Pal/-
 =  Lichadidae
 Familia: Callirhipidae Emden, 1924 [ ... R] -/-
 Superfamilia: Cebrionoidea [ ... ] Pal/
 =  Elateroidea
 Familia: Artematopodidae Lacordaire, 1857 [Pg23, BalJ ... R] Pal/Rus
 =  Artematopidae
 =  Eurypogonidae
 Subfamilia: Electribiinae Crowson, 1973 [Pg23, BalJ ... R] Pal/?
 Subfamilia: Artematopinae Lacordaire, 1857 [Pg23, BalJ ... R] Pal/Rus
 Familia: Brachypsectridae LeConte et Horn, 1883 [Pg33 ... R] -/-
 Familia: Rhinorhipidae Lawrence, 1987 [ ... R] -/-
+Familia: Praelateriidae Dolin, 1973 [J1he ... J1he] Pal/Rus
 Familia: Elateridae Leach, 1815 [J1he ... R] Pal/Rus
 =  Dicronychidae
 =  Lissomidae
 Subfamilia: Tetralobinae Laporte, 1840 [ ... R] -/-
 Subfamilia: Thylacosterninae Fleutiaux, 1920 [ ... R] ?/?
 =  Balginae
 =  Soleniscinae
 Subfamilia: Lissominae Laporte, 1835 [Pg1dn ... R] Pal/Rus
 Subfamilia: Oxynopterinae Cande`ze, 1857 [N1aq ... R] ?/?
 Subfamilia: Semiotinae Jacobson, 1913 [Pg1 ... R] ?/?
 Subfamilia: Pityobiinae Hyslop, 1917 [ ... R] -/-
+Subfamilia: Protagrypninae Dolin, 1973 [J1he ... J3ox] Pal/?
 Subfamilia: Agripninae Cande`ze, 1857 [J3ox ... R] Pal/Rus
 =  Cavicoxumidae
 =  Pangauridae Gistel, 1856
 =  Adelocerinae
 =  Campyloxeninae
 =  Hemirhipinae
 =  Pyrophorinae
 Subfamilia: Pyrophorinae Cande`ze, 1863 [Pg22 ... R]
 Subfamilia: Melanactinae Cande`ze, 1857 [ ... R]
 Subfamilia: Hypnoidinae Schwarz, 1906 [Pg23 ... R] Pal/Rus
 =?  Hypolithinae
 Subfamilia: Denticollinae Stein et Weise, 1877 [J3ox, KarT ... R] Pal/Rus
 =  Athoinae
 =  Campylinae
 =  Crepidomeninae
 =  Melanactinae
 Subfamilia: Diminae Cande`ze, 1859 [ ... R] ?/?
 Subfamilia: Elaterinae Leach, 1815 [Pg2 ... R] Pal/Rus
 =  Abrastinae
 Subfamilia: Hemiopinae Feutiaux, 1941 [ ... R] ?/?
 =  Hemiopsinae
 Subfamilia: Pleonominae Semenov-Tian-Shanskii et Piatakov, 1936 [ ... R]
 Subfamilia: Physodactylinae Lacordaire, 1857 [ ... R] ?/?
 Subfamilia: Aplastinae Stibick, 1979 [ ... R] ?/?
 Subfamilia: Oestodinae Hyslop, 1917 [K12 ... R] ?/?
 Subfamilia: Melanotinae Cande`ze, 1859 [ ... R] Pal/Rus
 Subfamilia: Negastriinae Nakane et Kishii, 1956 [J3ox ... R] Pal/Rus
 Subfamilia: Cardiophorinae Cande`ze, 1860 [J3ox ... R] Pal/Rus
 Subfamilia: Eudicronychinae Girard, 1971 [ ... R] ?/?
 =  Dicronychinae Schwartz, 1897
 Subfamilia: Anischiinae Feutiaux, 1936 [ ... R] ?/?
 Subfamilia: Suprotelaterinae Feutiaux, 1920 [ ... R] ?/?
 Familia: Eucnemidae Eschscholtz, 1829 [Pg23 ... R] Pal/Rus
 =  Melasidae Fleming, 1821
 =  Perothopidae Lacordaire, 1857
 =  Phylloceridae Reitter, 1905
 Subfamilia: Perothopinae Lacordaire, 1857 [ ... R] -/-
 Subfamilia: Phyllocerinae Reitter, 1905 [ ... R] Pal/-
 Subfamilia: Pseudomeninae Muona, 1993 [ ... R] -/-
 Subfamilia: Palaeoxeninae Muona, 1993 [ ... R] -/-
 Subfamilia: Phlegoninae Muona, 1993 [ ... R] -/-
 Subfamilia: Melasinae Fleming, 1821 [Pg31, Flor ... R] Pal/Rus
 Subfamilia: Eucneminae Eschscholtz, 1829 [ ... R] Pal/Rus
 Subfamilia: Macraulacinae Fleutiaux, 1923 [Pg23, BalJ ... R] ?/?
 Familia: Cebrionidae Latreille, 1802 [ ... R] Pal/-
 =  Aplastinae
 =  Cebriognathinae
 =  Plastocerinae
 Familia: Cerophytidae Latreille, 1834 [K12 ... R] Pal/Rus
 Familia: Throscidae Laporte, 1840 [K12 ... R] Pal/Rus
 =  Trixagidae
 Familia: Plastoceridae Crowson, 1972 [ ... R] Pal/-
 Superfamilia: Cantharoidea [ ... ] Pal/Rus
 Familia: Omalisidae Lacordaire, 1857 [ ... R] ?/?
 =  Homalisidae
 =  Omalysidae
 Familia: Berendtimiridae Winkler, 1987 [Pg23, BalJ ... R] ?/?
 Familia: Lycidae Laporte, 1836 [Pg23 ... R] Pal/Rus
 Subfamilia: Lycinae Laporte, 1836 [ ... R] Pal/Rus
 Subfamilia: Leptolycinae Leng et Mutchler, 1922 [ ... R] ?/?
 =  Duliticolinae Mjoeberg, 1925
 Subfamilia: Ateliinae Kleine, 1928 [ ... R] ?/?
 Subfamilia: Metriorrhynchinae Kleine, 1926 [ ... R] ?/?
 Subfamilia: Erotinae Leconte, 1881 [Pg23 ... R] Pal/Rus
 Subfamilia: Calochrominae Lacordaire, 1857 [ ... R] Pal/Rus
 Familia: Drilidae Blanchard, 1845 [ ... R] Pal/Rus
 Familia: Telegeusidae Leng, 1920 [ ... R] -/-
 Familia: Lampyridae Latreille, 1817 [Pg23 ... R] Pal/Rus
 Subfamilia: Pterotinae LeConte, 1861 [ ... R] -/-
 Subfamilia: Cyphonocerinae Crowson, 1972 [ ... R] ?/-
 Subfamilia: Ototretadrilinae Crowson, 1972 [ ... R] ?/-
 Subfamilia: Ototretinae McDermott, 1964 [ ... R] -/-
 Subfamilia: Amydetinae Olivier, 1907 [ ... R] ?/?
 Subfamilia: Lampyrinae Latreille, 1817 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Luciolinae Lacordaire, 1857 [Pg23 ... R] Pal/Rus
 Subfamilia: Photurinae Lacordaire, 1857 [ ... R] ?/?
 Familia: Omethidae LeConte, 1861 [ ... R] Pal/-
 Subfamilia: Driloniinae Crowson, 1972 [ ... R] -/-
 Subfamilia: Matheteinae LeConte, 1861 [ ... R] -/-
 Subfamilia: Omethinae LeConte, 1861 [ ... R] Pal/-
 Familia: Phengodidae LeConte, 1861 [ ... R] -/-
 Subfamilia: Rhagophthalminae Olivier, 1907 [ ... R] -/-
 Familia: Phengodidae LeConte, 1861 [ ... R] -/-
 Familia: Cantharidae Latreille, 1802 [Pg23 ... R] Pal/Rus
 =  Chaulignathidae
 =  Telephoridae
 Subfamilia: Cantharinae Latreille, 1802 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Dysmorphocerinae Brancucci, 1980 [ ... R] ?/?
 Subfamilia: Malthininae Kiesenwetter, 1852 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Chaulignathinae LeConte, 1861 [Pg31, Flor ... R] ?/?
 Subfamilia: Silinae Mulsant, 1862 [ ... R] Pal/Rus
 Familia: Podabrocephalidae Pic, 1930 [ ... R] -/-
 Infraordo: Cucujiformia [ ... ] Pal/Rus
 =  Chrysomelomorpha
 Superfamilia: Derodontoidea LeConte, 1861 [ ... R] Pal/Rus
 Familia: Derodontidae LeConte, 1861 [ ... R] Pal/Rus
 =  Laricobiidae Mulsant et Rey, 1863-1864
 =  Peltasticidae LeConte, 1861
 Subfamilia: Peltasticinae LeConte, 1861 [ ... R] Pal/Rus
 Subfamilia: Derodontinae LeConte, 1861 [ ... R] Pal/Rus
 Subfamilia: Laricobiinae Mulsant et Rey, 1863-1864 [ ... R] Pal/Rus
 Superfamilia: Bostrichoidea [K12 ... R] Pal/Rus
 =  Dermestoidea
 Familia: Nosodendridae Erichson, 1846 [Pg22 ... R] Pal/Rus
 Familia: Dermestidae Latreille, 1807 [K12 ... R] Pal/Rus
 =  Thorictidae
 =  Thylodriinae
 Subfamilia: Dermestinae Latreille, 1807 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Attageninae Laporte, 1840 [Pg23 ... R] Pal/Rus
 Subfamilia: Megatominae Leach, 1815 [Pg23 ... R] Pal/Rus
 Subfamilia: Orphilinae Leconte, 1861 [Pg31 ... R] Pal/Rus
 Subfamilia: Trinodinae Casey, 1900 [Pg23 ... R] Pal/Rus
 Subfamilia: Egidyellinae ???? [ ... R] Pal/-
 Subfamilia: Marioutinae Jacobson, 1913 [ ... R] Pal/-
 Subfamilia: Thorictinae Agassiz, 1846 [ ... R] Pal/?
 Subfamilia: Thylodriinae Semenov-Tian-Shanskii, 1913 [ ... R] Pal/Rus
 Familia: Endecotomidae LeConte, 1861 [ ... R] Pal/Rus
 Familia: Bostrichidae Latreille, 1802 [Pg21, LonC ... R] Pal/Rus
 =  Bostrychidae
 =  Lyctidae
 =  Psoidae
 Subfamilia: Dysidinae Lesne, 1921 [ ... R] ?/?
 =  Apoleoninae
 Subfamilia: Polycaininae Lesne, 1896 [ ... R] ?/?
 Subfamilia: Psoinae Blanchard, 1851 [ ... R] Pal/Rus
 Subfamilia: Dinoderinae C.G. Thomson, 1863 [Pg31 ... R] ?/?
 Subfamilia: Lyctinae Billberg, 1820 [Pg23 ... R] Pal/Rus
 Subfamilia: Euderinae Lesne, 1834 [ ... R] -/-
 Subfamilia: Bostrichinae Latreille, 1802 [Pg23 ... R] Pal/Rus
 Familia: Anobiidae Stephens, 1830 [K12 ... R] Pal/Rus
 =  Ectrephidae
 =  Gnostidae
 =  Ptinidae Latreille, 1802
 Subfamilia: Eucradinae LeConte, 1861 [Pg23 ... R] Pal/Rus
 =  Hedobiinae Mulsant et Rey, 1868
 Subfamilia: Ptininae Latreille, 1802 [Pg23 ... R] Pal/Rus
 =  Gibbiinae Mulsant et Rey, 1868
 Subfamilia: Ptilininae Shuckard, 1840 [Pg23 ... R] Pal/Rus
 Subfamilia: Dryophilinae LeConte, 1861 [ ... R] Pal/Rus
 Subfamilia: Alvarenganellinae Viana et Martinez, 1971 [ ... R] -/-
 Subfamilia: Ernobiinae Pic, 1912 [ ... R] Pal/Rus
 =  Cerocosminae Pic, 1912
 =  Cosmocerinae Solier, 1849
 Subfamilia: Xyletininae Gistel, 1856 [Pg23 ... R] Pal/Rus
 Subfamilia: Dorcatominae C.G. Thomson, 1859 [Pg23 ... R] Pal/Rus
 =  Dorkatominae
 Subfamilia: Mesocoelopodinae Mulsant et Rey, 1864 [Pg23 ... R] Pal/Rus
 =  Mesothinae Portevin, 1931
 =  Tricoryninae White, 1971
 Familia: Jacobsoniidae Heller, 1926 [ ... ]
 =  Sarothriidae Crowson, 1955
 =  Derolathrinae Sen Gupta, 1979
 Superfamilia: Lymexyloidea Fleming, 1921 [ ... ] Pal/Rus
 Familia: Lymexylidae Fleming, 1921 [Pg23 ... R] Pal/Rus
 =  Atractoceridae Laporte, 1840
 =  Lymexylonidae
 Subfamilia: Lymexylinae Fleming, 1921 [Pg23 ... R] Pal/Rus
 Subfamilia: Hylecoetinae Gistel, 1856 [N13 ... R] Pal/Rus
 Subfamilia: Melittommatinae Wheeler, 1986 [ ... R] -/-
 Superfamilia: Cleroidea Latreille, 1802 [J1to ... R] Pal/Rus
 Familia: Phloiophilidae Kisenwetter, 1863 [ ... R] Pal/?
 =  Phloeophilidae Kisenwetter, 1863
 Familia: Peltidae Kirby, 1837 [J1to ... R] Pal/Rus
 =  Larinotidae Slipinski, 1992
 =  Ostomidae Reitter, 1882
 Familia: Lophocateridae [ ... ] Pal/Rus
+Subfamilia: Meligethiellinae Kirejtshuk et Ponomarenko, 1990 [J31 ... ???] Pal/?
 Subfamilia: Protopeltinae Crowson, 1966 [ ... R] -/-
 Subfamilia: Larinotinae Slipinski, 1992 [ ... R] -/-
 Subfamilia: Rentoniinae Crowson, 1966 [ ... R] -/-
 Subfamilia: Decamerinae Crowson, 1964 [ ... R] -/-
 Subfamilia: Peltinae Latreille, 1807 [Pg23 ... R] Pal/Rus
 =  Ostominae
 Subfamilia: Lophocaterinae Crowson, 1964 [Pg23 ... R] Pal/Rus
 Familia: Trogossitidae Latreille, 1802 [ ... R] Pal/Rus
 =  Nemosomatidae Leach, 1815
 =  Nemosomidae
 =  Temnochilidae
 =  Tenebroididae
 Subfamilia: Gymnochilinae Lacordaire, 1854 [Pg23 ... R] Pal/Rus
 =  Calytinae Reitter, 1922
 Subfamilia: Egoliinae Lacordaire, 1854 [ ... R] ?/?
 Subfamilia: Trogossitinae Latreille, 1802 [Pg1d ... R] Pal/Rus
 =  Nemosominae
 =  Temnochilinae
 =  Tenebroidinae
 Familia: Chaetosomatidae Crowson, 1955 [ ... R] Pal/Rus
 Familia: Cleridae Latreille, 1802 [Pg13 ... R] Pal/Rus
 =  Corynetidae Laporte, 1836
 =  Korynetidae
 =  Thanerocleridae Chapin, 1924
 Subfamilia: Thaneroclerinae Chapin, 1924 [ ... R] ?/?
 Subfamilia: Tillinae Leach, 1815 [Pg23 ... R] Pal/Rus
 Subfamilia: Hydnocerinae Spinola, 1844 [Pg23 ... R] ?/?
 =  Phyllobaeninae Lacordaire, 1857
 Subfamilia: Clerinae Latreille, 1802 [Pg13 ... R] Pal/Rus
 =  Cleropiestinae Winkler, 1980
 =  Dieropsinae Winkler, 1964
 =  Priocerinae
 Subfamilia: Epiphleinae Kuwert, 1893 [ ... R] -/-
 Subfamilia: Tarsosteninae Jacquelin du Val, 1860 [Pg23 ... R] -/-
 Subfamilia: Korynetinae Laporte, 1836 [Pg23 ... R] Pal/Rus
 =  Corynetinae
 Familia: Acanthocnemidae Crowson, 1964 [ ... R] -/-
 Familia: Phycosecidae Crowson, 1952 [ ... R] -/-
 Familia: Prionoceridae Lacordaire, 1857 [ ... R] Pal/?
 Familia: Melyridae Leach, 1815 [ ... ] Pal/Rus
 =  Attalomenidae Majer, 1994
 =  Carphuridae
 =  Dasytidae Laporte, 1840
 =  Malachiidae Fleming, 1821
 =  Rhadalidae LeConte, 1861
 Subfamilia: Melyrinae Leach, 1815 [Pg23 ... R] Pal/Rus
 =  Attalomenidae Majer, 1994
 Subfamilia: Rhadalinae LeConte, 1861 [Pg23 ... R] Pal/?
 =  Aplocneminae
 =  Haplocneminae
 Subfamilia: Gietellinae Constantin de Minier, 1987 [ ... R] Pal/-
 Subfamilia: Dasytinae Laporte, 1840 [Pg23 ... R] Pal/Rus
 Subfamilia: Malachiinae Fleming, 1821 [ ... R] Pal/Rus
 Familia: Mauroniscidae Majer, 1994 [ ... R] -/-
 Superfamilia: Cucujoidea [ ... ] Pal/Rus
 =  Clavicornia
+Familia: Parandrexidae Kirejtshuk, 1993 [J21 ... J3ox] Pal/-
 Familia: Protocucujidae Crowson, 1954 [ ... R] -/-
 Familia: Sphindidae Jacquelin du Val, 1858 [Pg23 ... R] Pal/Rus
 =  Aspidiphoridae Kiesenwetter, 1877
 Subfamilia: Protosphindinae Sen Gupta et Crowson, 1979 [ ... R] -/-
 Subfamilia: Odontosphindinae Sen Gupta et Crowson, 1979 [ ... R] Pal/-
 Subfamilia: Sphindiphorinae Sen Gupta et Crowson, 1979 [ ... R] -/-
 Subfamilia: Sphindinae Jacquelin du Val, 1858 [Pg23 ... R] Pal/Rus
 =  Aspidiphoridae Kiesenwetter, 1877
 =  Eurysphindinae
 Familia: Helotidae Reitter, 1876/Chapuis, 1876 [ ... R] Pal/Rus
 Familia: Nitidulidae Latreille, 1802 [K12 ... R] Pal/Rus
 =  Carpophilidae Murray, 1864
 =  Cybocephalidae Jacquelin du Val, 1858
 =  Ipidae Lacordaire, 1854
 =  Mystropidae Murray, 1864
 =  Strongylinae Murray, 1864
 Subfamilia: Maynipeplinae Kirejtshuk, 1998 [ ... R] -/-
 Subfamilia: Calonecrinae Kirejtshuk, 1982 [ ... R] -/-
 Subfamilia: Epuraeinae Kirejtshuk, 1986 [ ... R] Pal/Rus
 Subfamilia: Carpophilinae Erichson, 1842 [ ... R] Pal/Rus
 Subfamilia: Amphicrossinae Kirejtshuk, 1986 [ ... R] Pal/Rus
 Subfamilia: Meligethinae C.G. Thomson, 1859 [Pg23 ... R] Pal/Rus
 Subfamilia: Nitidulinae Latreille, 1802 [K2cm ... R] Pal/Rus
 =  Strongylinae Erichson, 1843
 Subfamilia: Cillaeinae Kirejtshuk et Audisio, 1986 [Pg31 ... R] Pal/Rus
 Subfamilia: Cryptarchinae C.G. Thomson, 1859 [Pg23 ... R] Pal/Rus
 =  Ipinae Erichson, 1843
 =  Ipsinae Sharp, 1891
 =  Pityophaginae Fauconnet, 1894
 Subfamilia: Cybocephalinae Jacquelin du Val, 1858 [Pg23 ... R] Pal/Rus
 Familia: Boganiidae Sen Gupta et Crowson, 1966 [ ... R] -/-
 Subfamilia: Paracucujinae Endroedy-Younga et Crowson, 1986 [ ... R] -/-
 Subfamilia: Boganiinae Sen Gupta et Crowson, 1966 [ ... R] -/-
 Familia: Monotomidae Laporte, 1840 [Pg23 ... R] Pal/Rus
 =  Rhizophagidae Redtenbacher, 1845
 Subfamilia: Rhizophaginae Redtenbacher, 1845 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Monotominae [ ... ] Pal/Rus
 =  Lenacinae
 =  Thioninae
 Familia: Kateretidae Erichson, 1844 [Pg31 ... R] Pal/Rus
 =  Brachypteridae Erichson, 1845
 Familia: Smicripidae Horn, 1879 [ ... R] -/-
 Familia: Phloeostichidae Reitter, 1911 [ ... R] Pal/Rus
 Subfamilia: Phloestichinae Reitter, 1911 [ ... R] Pal/Rus
 Subfamilia: Hymaeinae Sen Gupta et Crowson, 1966 [ ... R] -/-
 Subfamilia: Agapythinae Sen Gupta et Crowson, 1966 [ ... R] -/-
 Subfamilia: Myraboliinae Lawrence et Britton, 1991 [ ... R] -/-
 Subfamilia: Priasilphinae Crowson, 1973 [ ... R] -/-
 Subfamilia: Tasmosalpinginae Lawrence et Britton, 1991 [ ... R] -/-
 Familia: Silvanidae Kirby, 1837 [Pg23 ... R] Pal/Rus
 Subfamilia: Brontinae Erichson, 1845/Blanchard, 1845 [ ... R] Pal/Rus
 =  Cryptamorphinae,
 =  Hyliotinae Reitter, 1879
 =  Psammoecinae
 =  Uleiotinae Gistel, 1856
 Subfamilia: Silvaninae Kirby, 1837 [Pg23 ... R] Pal/Rus
 Familia: Passandridae Erichson, 1845/Blanchard, 1845 [Pg23 ... R] Pal/?
 =  Catogenidae
 Familia: Cucujidae [ ... R] Pal/Rus
 Familia: Laemophloeidae Ganglbauer, 1899 [Pg22 ... R] Pal/Rus
 Familia: Propalticidae Crowson, 1952 [N2 ... R] -/-
 Familia: Phalacridae Leach, 1815 [Pg23 ... R] Pal/Rus
 =  Phaenocephalidae Matthews, 1899
 Subfamilia: Phaenocephalinae Matthews, 1899 [ ... R] Pal/-
 Subfamilia: Phalacrinae Leach, 1815 [Pg23 ... R] Pal/Rus
 Familia: Hobartiidae Sen Gupta et Crowson, 1966 [ ... ] -/-
 Familia: Cavognathidae Sen Gupta et Crowson, 1966 [ ... ] -/-
 Familia: Cryptophagidae Kirby, 1837 [K2sn ... R] Pal/Rus
 =  Catopochrotidae
 =  Hypocopridae Reitter, 1879
 Subfamilia: Hypocoprinae Reitter, 1879 [ ... R] Pal/Rus
 Subfamilia: Cryptophaginae Kirby, 1837 [Pg13 ... R] Pal/Rus
 =  Telmatophilinae
 Subfamilia: Atomariinae LeConte, 1861 [K2sn ... R] Pal/Rus
 =  Ephisteminae
 Familia: Lamingtoniidae Sen Gupta et Crowson, 1966 [ ... R] -/-
 Familia: Languriidae Crotch, 1873 [K1ap ... R] Pal/Rus
 Familia: Cryptophilidae Casey, 1900 [ ... R] Pal/Rus
 Subfamilia: Xenoscelinae Ganglbauer, 1899 [K1ap, Leba ... R] ?/?
 =  Loberinae
 =  Pharaxonotinae
 Subfamilia: Setariolinae Crowson, 1952 [ ... R] Pal/?
 =  Setariinae
 Subfamilia: Languriinae Crotch, 1873 [ ... R] Pal/Rus
 =  Cladoxeninae
 Subfamilia: Cryptophilinae Casey, 1900 [Pg23 ... R] Pal/Rus
 Subfamilia: Toraminae Sen Gupta, 1967 [ ... R] ?/?
 Familia: Erotylidae Latreille, 1802 [Pg23 ... R] Pal/Rus
 Familia: Dacnidae Gistel, 1856 [ ... ] Pal/Rus
 Subfamilia: Dacninae Gistel, 1856 [ ... R] Pal/Rus
 Subfamilia: Megalodacninae Sen Gupta, 1969 [ ... R] Pal/Rus
 Subfamilia: Encaustinae Crotch, 1876 [ ... R] ?/?
 Subfamilia: Tritominae Curtis, 1834 [Pg31 ... R] Pal/Rus
 =  Triplacinae Imhoff, 1856
 Subfamilia: Erotylinae Latreille, 1802 [ ... R] Pal/Rus
 Familia: Byturidae Jacquelin du Val, 1858 [K11 ... R] Pal/Rus
 Subfamilia: Platydascilinae Pic, 1914 [ ... R] ?/?
 Subfamilia: Byturinae Jacquelin du Val, 1858 [K11 ... R] Pal/Rus
 Familia: Biphyllidae LeConte, 1861 [ ... R] Pal/Rus
 =  Diphillidae
 Familia: Bothrideridae Erichson, 1845 [ ... R] Pal/Rus
 =  Anommatidae Ganglbauer, 1899
 Subfamilia: Teredinae Seidlitz, 1888 [ ... R] Pal/Rus
 Subfamilia: Xylariophilinae Pal et Lawrence, 1896 [ ... R] ?/?
 Subfamilia: Anommatinae Ganglbauer, 1899 [ ... R] ?/?
 Subfamilia: Bothriderinae Erichson, 1845 [Pg23 ... R] Pal/Rus
 =  Deretaphrinae
 Familia: Cerylonidae Billberg, 1820 [Pg23 ... R] Pal/Rus
 =  Aculagnathidae
 =  Cerylidae Billberg, 1820
 =  Dolosidae
 =  Euxestidae Grouvelle, 1908
 =  Murmidiidae Jacquelin du Val, 1858 (?)
 Subfamilia: Euxestinae Grouvelle, 1908 [ ... R] ?/?
 =  Metaceryloninae
 Subfamilia: Loeblioliryloninae Slipinski, 1990 [ ... R] ?/?
 Subfamilia: Ostomopsinae Sen Gupta et Crowson, 1973 [ ... R] ?/?
 Subfamilia: Murmidiinae Jacquelin du Val, 1858 [ ... R] Pal/Rus
 Subfamilia: Ceryloninae Billberg, 1820 [ ... R] Pal/Rus
 =  Aculagnathidae
 =  Lapethinae
 Familia: Alexiidae Imhoff, 1856 [ ... R] Pal/-
 =  Spharosomatidae Ganglbauer, 1899
 Familia: Discolomatidae Horn, 1878 [ ... R] Pal/?
 =  Aphanocephalidae Grouvelle, 1912
 =  Discolomidae
 =  Notiophygidae Jacobson, 1915
 Subfamilia: Notiophiginae Jacobson, 1915 [ ... R] -/-
 Subfamilia: Discolomatinae Horn, 1878 [ ... R] -/-
 Subfamilia: Apanocephalinae Grouvelle, 1912 [ ... R] Pal/?
 Subfamilia: Cephalophaninae John, 1954 [ ... R] -/-
 Subfamilia: Pondonatinae John, 1954 [ ... R] -/-
 Familia: Endomychidae Leach, 1815 [Pg23 ... R] Pal/Rus
 =  Cerasommatidae Bre'thes, 1925
 =  Merophysiidae Seidlitz, 1872
 =  Mycetaeidae Jacquelin du Val, 1858
 Subfamilia: Merophysiinae Seidlitz, 1872 [ ... R] Pal/Rus
 Subfamilia: Holoparamecinae Seidlitz, 1888 [Pg23 ... R] Pal/Rus
 Subfamilia: Leiestinae C.G. Thomson, 1863 [ ... R] Pal/Rus
 Subfamilia: Eupsilobiinae Casey, 1895 [ ... R] Pal/-
 =  Cerasommatidae Bre'thes, 1925
 =  Eidoreinae Sasaji, 1986
 Subfamilia: Endomychinae Leach, 1815 [ ... R] Pal/Rus
 Subfamilia: Epipocinae Gorham, 1873 [ ... R] Pal/Rus
 Subfamilia: Stenotarsinae Chapuis, 1876 [ ... ]
 Subfamilia: Lycoperdininae Redtenbacher, 1844 [Pg23, BalJ ... R] Pal/Rus
 =  Eumorphinae Gistel, 1856
 Subfamilia: Mycetaeinae Jacquelin du Val, 1858 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Anamorphinae Strohecker, 1953 [Pg23, BalJ ... R] ?/?
 =  Mychotheninae Sasaji, 1978
 Subfamilia: Pleganophorinae Jacquelin du Val, 1858 [Pg23 ... R] ?/?
 =  Trochoideinae Chapuis, 1876
 Subfamilia: Xenomycetinae Strohecker, 1962 [ ... R] -/-
 Familia: Coccinellidae Latreille, 1807 [K2cp ... R] Pal/Rus
 =  Epilachnidae
 Subfamilia: Coccidulinae Mulsant, 1846 [ ... ] Pal/Rus
 =  Tetrabrachinae
 Subfamilia: Coccinellinae Latreille, 1807 [Pg31, Flor ... R] Pal/Rus
 Subfamilia: Sticholotidinae Weise, 1901 [Pg23, BalJ ... R] Pal/Rus
 =  Pharinae Casey, 1899
 Subfamilia: Chilocorinae Mulsant, 1846 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Scymninae Mulsant, 1846 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Epilachninae Mulsant, 1846 [ ... R] Pal/Rus
 Familia: Corylophidae LeConte, 1852 [Pg23 ... R] Pal/Rus
 =  Orthoperidae
 Subfamilia: Peltinodinae Paulian, 1950 [ ... R] Pal/-
 Subfamilia: Corylophinae LeConte, 1852 [ ... R] Pal/Rus
 =  Orthoperinae
 Subfamilia: Sericoderinae Matthews, 1888 [ ... R] Pal/Rus
 Subfamilia: Parmulinae Poey, 1854 [ ... R] Pal/Rus
 =  Arthrolipinae
 =  Saciinae
 Familia: Latridiidae Erichson, 1842 [K2sn ... R] Pal/Rus
 =  Lathridiidae
 Subfamilia: Latridiinae Erichson, 1842 [Pg23 ... R] Pal/Rus
 Subfamilia: Corticariinae Curtis, 1829 [K2sn ... R] Pal/Rus
 Superfamilia: Tenebrionoidea Latreille, 1802 [J3ox ... R] Pal/Rus
 =  Heteromera
 Familia: Mycetophagidae Leach, 1815 [Pg23 ... R] Pal/Rus
 =  Tritomidae Crotch, 1873
 Subfamilia: Bergininae Leng, 1920 [ ... R] Pal/?
 Subfamilia: Esarcinae Reitter, 1882 [ ... R] Pal/?
 Subfamilia: Mycetophaginae Leach, 1815 [Pg23 ... R] Pal/Rus
 Familia: Archaeocrypticidae Kaszab, 1964 [ ... R] -/-
 Familia: Pterogeniidae Crowson, 1953 [ ... ]
 Familia: Ciidae Leach, 1819 [Pg23 ... R] Pal/Rus
 =  Cisidae
 =  Octotemnidae
 =  Orophidae
 Subfamilia: Sphindociinae Lawrence, 1974 [ ... R] -/-
 Subfamilia: Ciinae Leach, 1819 [Pg23 ... R] Pal/Rus
 Familia: Tetratomidae Billberg, 1820 [ ... R] Pal/Rus
 Subfamilia: Piseninae Miyatake, 1960 [ ... R] Pal/Rus
 Subfamilia: Tetratominae Billberg, 1820 [ ... R] Pal/Rus
 Subfamilia: Penthinae Lacordaire, 1859 [ ... R] Pal/?
 Familia: Melandryidae Leach, 1815 [K2sn ... R] Pal/Rus
 =  Serropalpidae
 Subfamilia: Hallomeninae Mulsant, 1856/Gistel, 1856 [K2sn, Jant ... R] Pal/Rus
 Subfamilia: Eustrophinae Gistel, 1856 [ ... R] ?/?
 Subfamilia: Melandryinae Leach, 1815 [K2sn, Jant ... R] Pal/Rus
 =  Dircaeinae
 =  Hypulinae
 =  Orchesiinae
 Subfamilia: Osphyinae Mulsant, 1856 [Pg23, BalJ ... R] Pal/Rus
 Familia: Mordellidae Latreille, 1802 [J3ox ... R] Pal/Rus
 =  Liaoximordellidae Wang, 1993
 =  Praemordellidae Schegoleva-Barovskaya, 1929
 Subfamilia: Ctenidiinae Franciscolo, 1951 [ ... R] -/-
 Subfamilia: Mordellinae Latreille, 1802 [J3ox ... R] Pal/Rus
 =  Praemordellinae Schegoleva-Barovskaya, 1929
 Familia: Scraptiidae Mulsant, 1856 [K11 ... R] Pal/Rus
 Familia: Anaspididae Mulsant, 1856 [ ... ] Pal/Rus
 Subfamilia: Scraptiinae Mulsant, 1856 [K11 ... R] Pal/Rus
 Subfamilia: Anaspidinae Mulsant, 1856 [ ... R] Pal/Rus
 Familia: Rhipiphoridae Harold et Gemminger, 1870 [Pg23 ... R] Pal/Rus
 Subfamilia: Pelecotominae Seidlitz, 1875 [Pg23 ... R] Pal/Rus
 Subfamilia: Micholaeminae Vianna, 1971 [ ... R] -/-
 Subfamilia: Ptilophorinae Gerstaecker, 1855 [ ... R] Pal/?
 =  Evaniocerinae
 Subfamilia: Hemirhipidiinae Heller, 1921 [ ... R] -/-
 =  Nephritinae Selander, 1957
 Subfamilia: Rhipidiinae Gerstaecker, 1855 [Pg23 ... R] Pal/Rus
 Subfamilia: Rhipiphorinae Gemminger et Harold, 1870 [Pg23 ... R] Pal/Rus
 Familia: Zopheridae Solier, 1834 [Pg23 ... R] Pal/Rus
 =  Colydiidae Erichson, 1842
 =  Monommatidae Blanchard, 1845
 =  Monommidae
 Subfamilia: Colydiinae Erichson, 1842 [Pg23 ... R] Pal/Rus
 Subfamilia: Pycnomerinae Erichson, 1845 [ ... R] ?/?
 Subfamilia: Zopherinae Solier, 1834 [ ... R] Pal/Rus
 =  Monommatinae Blanchard, 1845
 =  Monomminae
 =  Nosoderminae
 =  Usechinae Horn, 1867
 Familia: Ulodidae Pascoe, 1869 [ ... R] -/-
 =  Mericidae Crowson, 1953
 Familia: Perimylopidae St.George, 1939 [ ... R] -/-
 Familia: Chalcodryidae Watt, 1974 [ ... R] -/-
 Familia: Trachelostenidae Lacordaire, 1859 [ ... ] -/-
 Familia: Tenebrionidae Latreille, 1802 [J3ox ... R] Pal/Rus
 =  Alleculidae Laporte, 1840
 =  Cossiphodidae Wasmann, 1899
 =  Lagriidae Latreille, 1825
 =  Nilionidae Lacordaire, 1859
 =  Petriidae
 =  Rhysopaussidae
 =  Tenthiriidae
 Subfamilia: Lagriinae Latreille, 1825 [Pg23, BalJ ... R] Pal/Rus
 =  Lachninae Billberg, 1820
 =  Pycnocerinae
 Subfamilia: Cossyphinae Latreille, 1802 [ ... ]
 Subfamilia: Nilioninae Lacordaire, 1859 [ ... ]
 Subfamilia: Phrenapatinae Solier, 1834 [ ... R] ?/?
 Subfamilia: Zolodininae Watt, 1974 [ ... R] -/-
 Subfamilia: Pimeliinae Latreille, 1802 [Pg31, Flor ... R] Pal/Rus
 =  Tenthyriinae)
 Subfamilia: Tenebrioninae Latreille, 1802 [Pg23, BalJ ... R] Pal/Rus
 =  Toxicinae
 Subfamilia: Alleculinae Laporte, 1840 [J3ox, KarT ... R] Pal/Rus
 =  Cteniopodidae Solier, 1835
 =  Cistelinae Latreille, 1802
 =  Omophlinae
 =  Xystropodinae Solier, 1835
 Subfamilia: Diaperinae Latreille, 1802 [ ... R] Pal/Rus
 Subfamilia: Coelometopinae Lacordaire, 1859 [ ... R] ?/?
 =  Cnodalinae Gistel, 1856
 =  Stenochiinae Kirby, 1837
 Subfamilia: Cossyphodinae Wasmann, 1899 [ ... R] Pal/-
 Familia: Prostomidae C.G. Thomson, 1859 [ ... R] Pal/Rus
 Familia: Synchroidae Lacordaire, 1859 [Pg31, Flor ... R] ?/?
 Familia: Oedemeridae Latreille, 1810 [Pg23 ... R] Pal/Rus
 Subfamilia: Calopodinae Costa, 1852 [ ... R] Pal/Rus
 Subfamilia: Oedemerinae Latreille, 1810 [Pg23 ... R] Pal/Rus
 Familia: Stenotrachelidae C.G. Thomson, 1859 [ ... R] Pal/Rus
 =  Cephaloidae LeConte, 1862
 =  Nematoplidae LeConte, 1862
 Subfamilia: Stenotrachelinae C.G. Thomson, 1859 [ ... R] Pal/?
 Subfamilia: Nematoplinae LeConte, 1862 [ ... R] Pal/?
 Subfamilia: Cephaloinae LeConte, 1862 [ ... R] Pal/Rus
 Subfamilia: Stolinae Nikitsky, 1985 [ ... R] Pal/-
 Familia: Meloidae Gyllenhal, 1810 [Pg13 ... R] Pal/Rus
 =  Cantharidae Latreille, 1802
 =  Horiidae Latreille, 1802
 =  Tetraonychidae
 Subfamilia: Eleticinae Wellman, 1910 [ ... R] ?/?
 Subfamilia: Meloinae Gyllenhal, 1810 [Pg13 ... R] Pal/Rus
 =  Lyttinae
 =  Mylabrinae
 Subfamilia: Nemognathinae Laporte, 1840 [Pg31 ... R] Pal/Rus
 =  Horiinae
 =  Zonitidinae Mulsant, 1857
 Familia: Mycteridae Blanchard, 1845 [Pg23 ... R] Pal/Rus
 =  Hemipeplidae Lacordaire, 1854
 =  Rhinomaceridae Fleming, 1821
 Subfamilia: Mycterinae Blanchard, 1845 [N13 ... R] Pal/Rus
 Subfamilia: Lacconotinae LeConte, 1862 [ ... R] -/-
 =  Euryphinae J. Thomson, 1860
 Subfamilia: Hemipeplinae Lacordaire, 1854 [Pg23, BalJ ... R] ?/?
 Familia: Boridae C.G. Thomson, 1859 [ ... R] Pal/Rus
 Subfamilia: Borinae C.G. Thomson, 1859 [ ... R] Pal/Rus
 Subfamilia: Synercticinae Lawrence et Pollock, 1994 [ ... R] -/-
 Familia: Trictenotomidae Blanchard, 1845 [ ... R] Pal/-
 Familia: Pythidae Solier, 1834 [K2cm ... R] Pal/Rus
 =  Osphyoplessidae Reitter, 1917
 =?  Ischyomiidae Champion, 1886
 Familia: Pyrochroidae Latreille, 1807 [Pg23 ... R] Pal/Rus
 =  Cononotidae LeConte, 1862
 =  Pedilidae Lacordaire, 1859
 =  Pilipalpidae Abdullah, 1964
 =  Trictenotomidae Crowson, 1980
 Subfamilia: Tydessinae Nikitsky, 1986 [ ... R] Pal/Rus
 Subfamilia: Pilipalpinae Abdullah, 1964 [ ... R] -/-
 Subfamilia: Pedilinae Lacordaire, 1859 [ ... R] Pal/Rus
 Subfamilia: Pyrochroinae Latrielle, 1807 [ ... R] Pal/Rus
 Subfamilia: Agnathinae Lacordaire, 1859 [ ... R] Pal/?
 =  Cononotinae
 Familia: Salpingidae Leach, 1815 [Pg23 ... R] Pal/Rus
 =  Aegialitidae LeConte, 1862
 =  Dacoderidae LeConte, 1862
 =  Elacatidae Chapin, 1923
 =  Eurystethidae Seidlitz, 1916
 =  Inopeplidae Grouvele, 1908
 =  Othniidae LeConte, 1861
 =  Tretothoracidae
 Subfamilia: Othniinae LeConte, 1861 [ ... R] Pal/Rus
 =  Elacatinae
 Subfamilia: Prostominiinae Grouvelle, 1914 [ ... R] ?/?
 =  Trogocryptinae Crowson, 1953
 Subfamilia: Agleninae Horn, 1878 [ ... R] ?/?
 Subfamilia: Salpinginae Leach, 1815 [ ... R] Pal/Rus
 Subfamilia: Inopeplinae Grouvelle, 1908 [ ... R] Pal/?
 Subfamilia: Aegialitinae LeConte, 1862 [ ... R] Pal/Rus
 =  Eurystethinae
 Subfamilia: Dacoderinae LeConte, 1862 [ ... R] -/-
 =  Tretothoracinae
 Familia: Anthicidae Latreille, 1819 [K2-Pg2 ... R] Pal/Rus
 =  Notoxidae Stephens, 1829
 Subfamilia: Eurygeniinae LeConte, 1862 [K2-Pg2 ... R] ?/?
 Subfamilia: Lagriodinae Abdullah et Abdullah, 1968 [ ... R] -/-
 Subfamilia: Afreminae Levey, 1985 [ ... R] -/-
 Subfamilia: Macratriinae LeConte, 1862 [Pg23, BalJ ... R] -/-
 Subfamilia: Steropinae Jacquelin du Val, 1863 [Pg23, BalJ ... R] ?/?
 Subfamilia: Ischaliinae Blair, 1920 [ ... R] ?/?
 Subfamilia: Corobaeninae Abdullah, 1969 [ ... R] -/-
 Subfamilia: Lemodinae Lawrence et Britton, 1991 [ ... R] -/-
 Subfamilia: Tomoderinae Bonadona, 1961 [Pg23, BalJ ... R] ?/?
 Subfamilia: Anthicinae Latreille, 1819 [Pg23, BalJ ... R] Pal/Rus
 =  Notoxinae
 Familia: Aderidae Winkler, 1927 [Pg23 ... R] Pal/Rus
 =  Circeidae Iablokoff-Khnzorian, 1961
 =  Euglenesidae
 =  Euglenidae Seidlitz, 1875
 =  Hylophilidae Pic, 1900
 =  Xylophilidae Shuckard, 1840
 Superfamilia: Chrysomeloidea Latreille, 18?? [ ... ] Pal/Rus
 Familia: Megalopodidae Latreille, 1802 [ ... R] Pal/Rus
 Subfamilia: Palophaginae Kuschel et May, 1990 [ ... R] -/-
 Subfamilia: Zeugophorinae Boeving et Craigheead, 1931 [Pg23 ... R] Pal/Rus
 Subfamilia: Megalopodinae Latreille, 1802 [ ... R] Pal/Rus
 Familia: Orsodacnidae C.G. Thomson, 1859 [ ... R] Pal/Rus
 Subfamilia: Orsodacninae C.G. Thomson, 1859 [ ... R] Pal/Rus
 Subfamilia: Aulacoscelidinae Chapius, 1874 [ ... R] -/-
 Familia: Chrysomelidae Latreille, 1802 [J31 ... R] Pal/Rus
 =  Sagridae Leach, 1815
 Familia: Bruchidae Latreille, 1802 [ ... ] Pal/Rus
 Subfamilia: Bruchinae Latreille, 1802 [Pg31 ... R] Pal/Rus
 =  Pachymerinae Bridwell, 1929
 =  Rhaebinae Chapuis, 1874
 Subfamilia: Hisperinae ??? [ ... R] Pal/Rus
 Subfamilia: Crysomelinae ??? [ ... R] Pal/Rus
 Subfamilia: Eumolpinae ??? [ ... R] Pal/Rus
 Subfamilia: Galerucinae Latreille, 1802 [Pg23 ... R] Pal/Rus
 =  Alticinae
 =  Halticinae Newman, 1834,
 Subfamilia: Donaciinae Kirby, 1837 [Pg1d ... R] Pal/Rus
 Subfamilia: Criocerinae Latreille, 1804 [Pg23 ... R] Pal/Rus
 Subfamilia: Lamprosomatinae Lacordaire, 1848 [ ... ]
 Subfamilia: Cryptocephalinae Gyllenhal, 1813 [Pg13 ... R] Pal/Rus
 =  Chlamisinae
 =  Clytrinae Kirby, 1837
 Familia: Cerambycidae Latreille, 1802 [K12 ... R] Pal/Rus
 =  Hypocephalidae Blanchard, 1845
 =  Parandridae Blanchard, 1845
 =  Spondylidae
 Subfamilia: Parandrinae Blanchard, 1845 [ ... R] Pal/Rus
 Subfamilia: Prioninae Latreille, 1802 [ ... R] Pal/Rus
 Subfamilia: Spondylidinae Audinet-Serville, 1832 [ ... R] Pal/Rus
 Subfamilia: Apatophyseinae Lacordaire, 1869 [ ... R] Pal/?
 Subfamilia: Necydalinae Latreille, 1825 [ ... R] ?/?
 Subfamilia: Lepturinae Latreille, 1802 [ ... R] Pal/Rus
 Subfamilia: Cerambycinae Latreille, 1802 [ ... R] Pal/Rus
 Subfamilia: Lamiinae Latreille, 1825 [ ... R] Pal/Rus
 Familia: Disteniidae J. Thomson, 1860 [ ... R] ?/?
 Familia: Vesperidae Mulsant, 1839 [ ... R] Pal/?
 =  Anoplodermatidae Gue'rin-Me'neville, 1840
 Subfamilia: Anoplodermatinae Gue'rin-Me'neville, 1840 [ ... R] ?/?
 Subfamilia: Philinae J. Thomson, 1860 [N12 ... R] ?/?
 =  Hypocephalinae Blanchard, 1845
 Familia: Oxypeltidae Lacordaire, 1869 [ ... R] -/-
 Infraordo: Curculioniformia Kirejtshuk, 1991 [ ... ] Pal/Rus
 Superfamilia: Curculionoidea [ ... ] Pal/Rus
 =  Rhynchophora Latreille, 1817
+Familia: Obrienidae Zherikhin et Gratshev, 1993 [T2 ... J13] Pal/?
+Subfamilia: Obrieninae Zherikhin et Gratshev, 1993 [??? ... ???] Pal/?
+Subfamilia: Kararhynchinae Zherikhin et Gratshev, 1993 [??? ... ???] Pal/?
+Familia: Ulyanidae Zherikhin, 1993 [K13 ... K13]
 Familia: Nemonychidae Bedel, 1882 [J31 ... R] Pal/Rus
 =  Eubelidae Arnoldi, 1977
 =  Rhinomaceridae Schoenherr, 1832
+Subfamilia: Eobelinae Arnoldi, 1977 [J31 ... ???] Pal/?
+Subfamilia: Brenthorrhininae Arnoldi, 1977 [??? ... ???] Pal/?
+Subfamilia: Nanophydinae Arnoldi, 1977 [??? ... ???] Pal/?
+Subfamilia: Oxycorynoidinae Arnoldi, 1977 [??? ... ???] Pal/?
 Subfamilia: Nemonychinae Bedel, 1882 [ ... R] Pal/Rus
 Subfamilia: Rhinorhynchinae Voss, 1922 [ ... R] -/-
 Subfamilia: Doydirhynchinae Pierce, 1916 [ ... R] Pal/-
 =  Cimberidinae Gozis, 1882
 =  Neocimberidinae
+Subfamilia: Slonikinae Zherikhin, 1977 [ ... ]
 Familia: Anthribidae Billberg, 1820 [K1 ... R] Pal/Rus
 =  Bruchelidae
 =  Platyrhynidae
 =  Platystomatidae
 =  Platystomidae
 =  Urodontidae
 Subfamilia: Urodontinae C.G. Thomson, 1859 [ ... R] Pal/Rus
 =  Bruchelinae
 Subfamilia: Anthribinae Billberg, 1820 [ ... R] Pal/Rus
 =  Brachytarsinae
 Subfamilia: Choraginae Kirby, 1819 [ ... R] ?/?
 =  Araeocerinae
+Familia: Eccortarthridae Arnoldi, 1977 [J?? ... ???] Pal/?
+Subfamilia: Eccortarthrinae Arnoldi, 1977 [J?? ... ???] Pal/?
+Subfamilia: Baissorhynchinae Zherikhin, 1993 [ ... ] Pal/?
 Familia: Belidae Schoenherr, 1826 [ ... R] ?/?
 Subfamilia: Belinae Schoenherr, 1826 [ ... R] ?/?
 Subfamilia: Pachyurinae Kuschel, 1959 [ ... R] -/-
 Familia: Oxycorynidae Schoenherr, 1840 [ ... R] ?/?
 Subfamilia: Oxycoryninae Schoenherr, 1840 [ ... R] ?/?
 Subfamilia: Afrocoryninae Voss, 1957 [ ... R] -/-
 Subfamilia: Aglycyderinae Wollaston, 1864 [ ... R] -/-
 =  Proterhininae
 Subfamilia: Allocoryninae Sharp, 1890 [ ... R] ?/?
 Subfamilia: Hispodinae Voss, 1957 [ ... R] ?/?
 Subfamilia: Metrioxeninae Voss, 1953 [ ... R] ?/?
 Familia: Rhynchitidae Gistel, 1865 [ ... R] Pal/Rus
 =  Pterocolidae Lacordaire, 1866
 Subfamilia: Rhynchitinae Gistel, 1865 [ ... R] Pal/Rus
 Subfamilia: Pterocolinae Lacordaire, 1866 [ ... R] ?/?
+Subfamilia: Isotheinae Scudder, 1893 [??? ... ???] ?/?
 Familia: Attelabidae Billberg, 1820 [K12 ... R] Pal/Rus
 =  Apoderidae Jekel, 1860
 Subfamilia: Attelabinae Billberg, 1820 [ ... R] Pal/Rus
 Subfamilia: Apoderinae Jekel, 1860 [ ... R] Pal/Rus
 Subfamilia: Archolabinae Voss, 1929 [ ... R] ?/?
 Familia: Brentidae Billberg, 1820 [Pg33 ... R] Pal/Rus
 =  Brenthidae
 =  Cyladidae
 =  Eurhynchidae
 Subfamilia: Brentinae Billberg, 1820 [ ... R] Pal/Rus
 Subfamilia: Antliarhininae Schoenherr, 1823 [ ... R] ?/?
 Subfamilia: Cyladinae [ ... R] ?/?
 Subfamilia: Cyphagoginae Kolbe, 1892 [ ... R] ?/?
 Subfamilia: Pholidochlamidinae Damoiseau, 1962 [ ... R] ?/?
 Subfamilia: Taphroderinae Lacordaire, 1866 [ ... R] ?/?
 Subfamilia: Trachelizinae Lacordaire, 1866 [ ... R] ?/?
 Subfamilia: Ulocerinae Schoenherr, 1823 [ ... R] ?/?
 Subfamilia: Eurhynchinae Lacordaire, 1863 [ ... R] ?/?
 Familia: Apionidae Schoenherr, 1823 [ ... R] Pal/Rus
 Subfamilia: Apioninae Schoenherr, 1823 [ ... R] Pal/Rus
 Subfamilia: Tanainae [ ... R] ?/?
 Subfamilia: Myrmacicelinae Zimmerman, 1994 [ ... R] -/-
 Subfamilia: Rhinorhynchidiinae Zimmerman, 1994 [ ... R] -/-
 Subfamilia: Nanophyinae Gistel, 1856 [ ... R] Pal/Rus
 Familia: Caridae Thomson, 1992 [ ... R] -/-
 Familia: Ithyceridae Schoenherr, 1823 [ ... R] -/-
 Familia: Curculionidae Latreille, 1802 [K13 ... R] Pal/Rus
 =  Calendridae
 =  Cossonidae Schoenherr, 1825
 =  Platypodidae
 =  Rhynchophoridae Schoenherr, 1833
 =  Scolytidae
 Subfamilia: Brachycerinae Billberg, 1820 [ ... R] Pal/Rus
 =  Alophinae LeConte, 1874
 =  Amycterinae G.R.Waterhouse, 1854
 =  Aterpinae Lacordaire, 1863
 =  Brachyderinae Schoenherr, 1826
 =  Byrsopinae Germar, 1829
 =  Cryptolaringinae Schalkwyk, 1966
 =  Cylindrorhininae
 =  Desmidophorinae Morimoto, 1962
 =  Diabathrariinae Lacordaire, 1863
 =  Dinomorphinae Lacordaire, 1863
 =  Ectemnorhininae Lacordaire, 1863
 =  Entiminae Schoenherr, 1823
 =  Eremninae
 =  Goniopterinae Lacordaire, 1863
 =  Hipporhininae
 =  Hyperinae Lacordaire, 1863
 =  Leptopiinae
 =  Microcerinae Lacordaire, 1863
 =  Ocladiinae Lacordaire, 1866
 =  Otiorhynchinae Schoenherr, 1826
 =  Pachyrhynchinae Schoenherr, 1826
 =  Rhadinosominae
 =  Rhyparosominae
 =  Rhytirhininae Lacordaire, 1863
 =  Somatodinae Lacordaire, 1863
 =  Tanyrhynchinae Schoenherr, 1826
 =  Thecesterninae Lacordaire, 1863
 =  Ulomascinae Lacordaire, 1866
 =  Viticinae Morimoto, 1983
 Subfamilia: Curculioninae Latreille, 1802 [ ... R] Pal/Rus
 =  Anoplinae Bedel, 1884
 =  Anthonominae C.G.Thomson, 1859
 =  Bagoinae C.G.Thomson, 1859
 =  Baridinae Schoenherr, 1836
 =  Camarotinae Schoenherr, 1833
 =  Campyloscelinae Schoenherr, 1845
 =  Ceratopodinae Lacordaire, 1863
 =  Ceutorhynchinae Gistel, 1856
 =  Cioninae Schoenherr, 1825
 =  Cleoninae Schoenherr, 1826
 =  Cryptorhynchinae Schoenherr, 1825 ?/?
 =  Erirhininae Schoenherr, 1825
 =  Eugnominae Lacordaire, 1863
 =  Tadiinae Zimmerman, 1993
 Subfamilia: Gymnetrinae [ ... ]
 =  Lixinae Schoenherr, 1823
 =  Magdalininae Pascoe, 1870
 =  Molytinae Schoenherr, 1823
 =  Nerthopinae Lacordaire, 1866
 =  Otidocephalinae Lacordaire, 1863
 Subfamilia: Prionomerinae [ ... ]
 =  Raymondionymidae Reitter, 1913
 =  Myrtonyminae Kuschel, 1990
 =  Pyropinae Lacordaire, 1866
 =  Rhyncheninae
 =  Tachigoninae
 =  Trigonocolinae Lacordaire, 1863
 =  Tychiinae C.G.Thomson, 1859
 =  Zygopinae Lacordaire, 1866
 Subfamilia: Brachyceropseinae Aurivillius, 1926 [ ... R] ?/?
 Subfamilia: Conoderinae Schoenherr, 1833 [ ... R] ?/?
 Subfamilia: Cyclominae Schoenherr, 1826 [ ... R] ?/?
 Subfamilia: Mesoptiliinae Lacordaire, 1863 [ ... R] ?/?
 Subfamilia: Orobitidinae C.G.Thomson, 1859 [ ... R] ?/?
 Subfamilia: Xiphaspidinae Marshall, 1920 [ ... R] ?/?
 Subfamilia: Dryophthorinae Schoenherr, 1825 [ ... R] ?/?
 =  Calandrinae
 =  Calendrinae
 =  Cryptodermatinae Bovie, 1908
 =  Orthognathinae Lacordaire, 1866
 =  Rhynchophorinae Schoenherr, 1833
 =  Stromboscerinae Lacordaire, 1866
 Subfamilia: Cossoninae Schoenherr, 1825 [ ... R] Pal/Rus
 Subfamilia: Scolytinae Latreille, 1804 [ ... R] Pal/Rus
 =  Coptonotinae
 =  Hylesininae
 =  Ipinae
 =  Tomicinae
 Subfamilia: Platypodinae Shuckard, 1840 [ ... R] Pal/Rus
 =  Chapuisiinae
 =  Crossotarsinae
 =  Diaporinae
 =  Periommatinae
 =  Platytarsilinae
 Subordo: Myxophaga Crowson, 1955 [ ... ] Pal/Rus
 Infraordo: Microsporomorpha ???Crotch, 1873 [ ... ] Pal/Rus
+Superfamilia: Asiocoleoidea Rohdendorf, 1961 [ ... ]
+Familia: Asiocoleidae Rohdendorf, 1961 [P2uf ... P2kz] Pal/?
+Familia: Tricoleidae Ponomarenko, 1969 [T2l ?T3r ... T3] Pal/?
+Superfamilia: Rhombocoleoidea Rohdendorf, 1961 [ ... ]
+Familia: Rhombocoleidae Rohdendorf, 1961 [P2uf ... P2t] Pal/?
+Superfamilia: Schizophoroidea Ponomarenko, 1968 [ ... ]
+Familia: Schizophoridae Ponomarenko, 1968 [P2kz ... K1 (?K2)] Pal/?
+Familia: Catiniidae Ponomarenko, 1968 [T2l ... J31] Pal/?
+Familia: Schizocoleidae Rohdendorf, 1961 [P2kz, Suri ... K1ap, Bon] Pal/?
 =  Curculiopseidae Martynov, 1937
 Superfamilia: Microsporoidea Crotch, 1873 [ ... R]
 Familia: Lepiceridae Hinton, 1936 [ ... R] -/-
 =  Cyathoceridae Scharp, 1882
 Familia: Microsporidae Crotch, 1873 [ ... R] Pal/Rus
 =  Sphaeriidae Erichson, 1845
 Familia: Hydroscaphidae LeConte, 1874 [ ... R] Pal/?
 Familia: Torridincolidae Steffan, 1964 [ ... R] Pal/?
 Subordo: Adephaga Schellenberg, 1806 [ ... ] Pal/Rus
 Infraordo: Haliplomorpha Brulle', 1835 [ ... ] Pal/Rus
 Superfamilia: Triaploidea Ponomarenko, 1977 [ ... ]
 Familia: Triaplidae Ponomarenko, 1977 [T2l ... T3c] Pal/?
 Superfamilia: Haliploidea [ ... ] Pal/Rus
 Familia: Haliplidae Brulle', 1835 [Kal3-cm1 ... R] Pal/Rus
 Infraordo: Carabomorpha [ ... ] Pal/Rus
 Superfamilia: Gyrinoidea ???Latreille, 1810 [ ... ] Pal/Rus
 Familia: Gyrinidae Latreille, 1810 [J11 ... R] Pal/Rus
 Subfamilia: Spanglerogyrininae Folkerts, 1979 [ ... R] -/-
 Subfamilia: Gyrininae Latreille, 1810 [J11 ... R] Pal/Rus
 =  Orectochilinae
 Superfamilia: Dytiscoidea Leach, 1815 [ ... ] Pal/Rus
 Familia: Colymbothetidae Ponomarenko, 1993 [T3r ... ???] Pal/?
 Familia: Hygrobiidae Regimbart, 1878 [N1aq ... R] Pal/?
 =  Pelobiidae
 Familia: Parahygrobiidae Ponomarenko, 1977 [J31 ... ???] Pal/?
 Familia: Coptoclavidae Ponomarenko, 1961 [J11 ... K1ap] Pal/?
 Subfamilia: Timarchopsinae [J11 ... K1ha] Pal/?
 =  Necronectinae Ponomarenko, 1977
 Subfamilia: Charonoscaphinae Ponomarenko, 1977 [J3ox, KarT ... ???] Pal/?
 Familia: Liadytidae Ponomarenko, 1977 [J11 ... J3] Pal/?
 Familia: Noteridae Thomson, 1860 [Pg12 ... R] Pal/Rus
 =  Phreatodytidae Ueno, 1957
 Familia: Noteridae Thomson, 1860 [Pg12 ... R] Pal/Rus
 Familia: Phreatodytidae Ueno, 1957 [ ... R] Pal/-
 Familia: Amphizoidae Lacordaire, 1854 [ ... R] Pal/?
 Familia: Dytiscidae Leach, 1815 [J31 ... R] Pal/Rus
 Subfamilia: Palaeogyrininae Hatch, 1927 [Pg12 ... N1aq, Rot] ?/?
 Subfamilia: Copelatinae Van den Branden, 1885 [N12, Stav ... R] Pal/Rus
 Subfamilia: Aubehydrinae Guignot, 1942 [ ... R] ?/?
 Subfamilia: Laccophilinae Gistel, 1856 [Pg22, GreR ... R] Pal/Rus
 Subfamilia: Hydroporinae Sharp, 1880 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Colymbetinae Erichson, 1837 [Pg23, BalJ ... R] Pal/Rus
 Subfamilia: Dytiscinae Leach, 1815 [Pg31, Flor ... R] Pal/Rus
 Superfamilia: Caraboidea ???Latreille, 1802 [T2l ... R] Pal/Rus
 Familia: Trachypachidae Thomson, 1857 [T2l ... R] Pal/Rus
 =  Leptopodocoleidae Hong, 1982
 Subfamilia: Trachypachinae Thomson, 1857 [T2l ... R] Pal/Rus
 Subfamilia: Eodromeinae Ponomarenko, 1977 [T2-3l-ca ... K1ap] Pal/?
 =  Leptopodocoleinae Hong, 1982
 Familia: Carabidae Latreille, 18O2 [J11 ... R] Pal/Rus
 =  Cicindelidae
 =  Paussidae
 Subfamilia: Protorabinae Ponomarenko, 1977 [J1he ... K1al] Pal/?
 Subfamilia: Gehringiinae Darlington, 1933 [ ... R] ?/?
 Subfamilia: Cicindelinae Latreille, 1802 [Pg22 ... R] Pal/Rus
 Subfamilia: Paussinae Latreille, 1807 [Pg23 ... R] Pal/Rus
 Subfamilia: Nototylinae Reichardt, 1977 [ ... R] ?/?
 Subfamilia: Cicindinae Baenninger, 1927 [ ... R] Pal/Rus
 Subfamilia: Metriinae Leconte, 18?? [Pg23, BalJR ... R]
 Subfamilia: Omophroninae Bonelli, 1810 [N13 ... R] Pal/Rus
 Subfamilia: Carabinae Latreille, 1802 [K2tu ... R] Pal/Rus
 Subfamilia: Hiletinae Schioedte, 1847 [ ... R] ?/?
 Subfamilia: Loricerinae Bonelli, 1810 [ ... R] ?/?
 Subfamilia: Elaphrinae Latreille, 1802 [ ... R] Pal/Rus
 Subfamilia: Migadopinae Chadoir, 1861 [ ... R] ?/?
 Subfamilia: Siagoninae Bonelli, 1810 [ ... R] ?/?
 Subfamilia: Scaritinae Bonelli, 1810 [ ... R] Pal/Rus
 Subfamilia: Trechinae Bonelli, 1810 [ ... R] Pal/Rus
 Subfamilia: Harpalinae Bonelli, 1810 [ ... R] Pal/Rus
 Subfamilia: Pseudomorphinae Newmann, 1842 [ ... R] -/-
 Subfamilia: Brachininae Bonelli, 1810 [ ... R] Pal/Rus
 Superfamilia: Rhysodoidea ???Laporte, 1840 [Pg33 ... R] Pal/Rus
 Familia: Rhysodidae Laporte, 1840 [Pg33 ... R] Pal/Rus