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Karen A. Ober, David R. Maddison. 2008.
Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene.
Journal of Insect Science, 2008. Vol. 8. Article 63. 32 pp.
Journal of Insect Science 8:63, available online: insectscience.org/8.63