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Bert Kohlmann, Angel Solis, Guillermo E. Alvarado. 2019.
Description of Onthophagus humboldti and Uroxys bonplandi, two new scarab beetles (Coleoptera, Scarabaeidae, Scarabaeinae) from Costa Rica, with notes on tropical mountain brachyptery and endemicity.
ZooKeys, 2019, 881: 23-51.
doi: 10.3897/zookeys.881.38026
http://zoobank.org/ADCA057E-3E1B-45D8-B5C9-665683F51B59
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