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Bert Kohlmann, Alfonsina Arriaga-Jiménez, Matthias Rös. 2018.
Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae).
ZooKeys, 2018, 743: 67-93.
https://doi.org/10.3897/zookeys.743.23029
http://zoobank.org/4D31DBBF-07B1-479C-9048-9C9207B5103D