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Wataru Ichiishi, Shinpei Shimada, Takashi Motobayashi, Hiroaki Abe. 2019.
Completely engaged three-dimensional mandibular gear-like structures in the adult horned beetles: reconsideration of bark-carving behaviors (Coleoptera, Scarabaeidae, Dynastinae).
ZooKeys, 2019, 813: 89-110.
doi: 10.3897/zookeys.813.29236
http://zoobank.org/84A4ADAA-4301-4890-BC45-04985BAFDBA2