Family Scarabaeidae Tribe Onthophagini (Scarabaeidae: Scarabaeinae) - Atlas of scarab beetles of Russia Onthophagus (Palaeonthophagus) gibbulus gibbulus (Pallas, 1781) 2.06.2007. Island Sàkhàlin, Listvånnichnîå. Male and Female. leg. Vårtyankin À.Â.