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Grebennikov, V.V., Ballerio, A., Ocampo, F. and Scholtz C.H. 2004.
Larvae of Ceratocanthidae and Hybosoridae (Coleoptera: Scarabaeoidea): study of morphology, phylogenetic analysis and evidence of paraphyly of Hybosoridae.
Systematic Entomology, 29: 524-543.